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Division of HaematologyJohns Hopkins University Medical SchoolBaltimore Maryland 3393_C000.fm Page vi Monday, November 19, 2007 1:58 PM... ADCC antigen-dependent cellular cytotoxicity AM

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CRC Press Taylor & Francis Group

6000 Broken Sound Parkway NW, Suite 300 Boca Raton, FL 33487-2742

© 2008 by Taylor & Francis Group, LLC CRC Press is an imprint of Taylor & Francis Group, an Informa business

No claim to original U.S Government works Printed in the United States of America on acid-free paper

10 9 8 7 6 5 4 3 2 1 International Standard Book Number-13: 978-0-8493-3393-4 (Hardcover) This book contains information obtained from authentic and highly regarded sources Reprinted material is quoted with permission, and sources are indicated A wide variety of references are listed Reasonable efforts have been made to publish reliable data and information, but the author and the publisher cannot assume responsibility for the validity of all materials or for the consequences of their use

Except as permitted under U.S Copyright Law, no part of this book may be reprinted, reproduced, transmitted, or lized in any form by any electronic, mechanical, or other means, now known or hereafter invented, including photocopy- ing, microfilming, and recording, or in any information storage or retrieval system, without written permission from the publishers.

uti-For permission to photocopy or use material electronically from this work, please access www.copyright.com (http:// www.copyright.com/) or contact the Copyright Clearance Center, Inc (CCC) 222 Rosewood Drive, Danvers, MA 01923, 978-750-8400 CCC is a not-for-profit organization that provides licenses and registration for a variety of users For orga- nizations that have been granted a photocopy license by the CCC, a separate system of payment has been arranged.

Trademark Notice: Product or corporate names may be trademarks or registered trademarks, and are used only for

identification and explanation without intent to infringe.

Library of Congress Cataloging-in-Publication Data

Desk reference for hematology / edited by N.K Shinton 2nd ed.

p ; cm.

Rev ed of: CRC desk reference for hematology c1998.

Includes bibliographical references and index.

ISBN 0-8493-3393-8 (alk paper)

1 Hematology Handbooks, manuals, etc 2 Blood Diseases Handbooks, manuals, etc 3

Blood Handbooks, manuals, etc I Shinton, N K (N Keith) II CRC desk reference for hematology

[DNLM: 1 Hematologic Diseases Handbooks 2 Hematology Handbooks WH 39 D459 2006]

RB145.C69 2006

Visit the Taylor & Francis Web site at http://www.taylorandfrancis.com and the CRC Press Web site at http://www.crcpress.com

3393_C000.fm Page ii Monday, November 19, 2007 1:58 PM

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CRC Press is an imprint of the

Taylor & Francis Group, an informa business

Boca Raton London New York

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The intention of the publishers, editor, and authors of this book is to provide a majorsource of easily obtained, reliable information on hematology in one book, with specifictopics arranged in alphabetical order As this is mainly based on well-established knowl-edge, it has not been closely referenced The references provided here have been confined

to guidelines, reviews, and recently published articles on topics that are either sial or for which opinion has yet to be consolidated In addition, there is a bibliography

controver-of recommended further reading

The information has been made concise by the use of extensive cross-referencing withinthe book, as indicated by words printed in boldface Words thus highlighted within anarticle indicate that further information on the subject is available in a separate articleidentified by the emboldened words

Common abbreviations are listed in a separate table at the front of the book and arewidely identified throughout the text Acronyms for combination therapies are listedunder cytotoxic agents Manuscript references are superscript numbers; in-text referencesare numbers in brackets [1]

The nomenclature regarding hematopoietic and lymphoid tissue is that used in theWorld Health Organization (WHO) Classification of Tumors of Hematopoietic and Lym-phoid Tissues, a summary of which is included as an Appendix Color photographs ofcells or tissues have not been included, as these are well presented in many easily obtain-able publications Likewise, details of laboratory technical procedures are not included

It is my great pleasure to acknowledge the help that I have received from the authorsand from colleagues, in particular Dr R A Shinton, senior lecturer in medicine, University

of Birmingham, Birmingham, U.K., for reading the manuscript and Mrs Anne Caine,administrator, Division of Clinical Sciences, Warwick Medical School, University of War-wick, Coventry, U.K., for help in data processing

N K Shinton

Editor3393_C000.fm Page iv Monday, November 19, 2007 1:58 PM

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The indications and dosage of all drugs in this book are those recommended in the medicalliterature and in conformance with general medical practice The drugs do not necessarilyhave specific approval by the U.S Food and Drug Administration or of the EuropeanPharmaceutical Commission, either for use in the disorders or in the dosages recom-mended The package insert for each drug should be consulted for indications, contrain-dications, dosage, and adverse drug reactions, particularly in respect to weight and age

in children It is advisable for those using a specific drug to be aware of any revisedrecommendations, especially when prescribing drugs that have recently become available.3393_C000.fm Page v Monday, November 19, 2007 1:58 PM

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Rajesh Chopra, Ph.D., F.R.C.P., F.R.C Path.

University College Hospital Medical School

London, U.K

Christopher D Fegan, M.D., F.R.C.P., F.R.C.Path.

Llandaff HospitalCardiff, Wales, U.K

Edward C Gordon-Smith, M.A., M.Sc., M.B., F.R.C.P., F.R.C.Path.

Robert A Kyle, M.D.

Department of Hematology and Internal Medicine

Mayo ClinicRochester, Minnesota

Edwin Massey, M.B., F.R.C.P., F.R.C.Path.

National Blood ServiceBristol, U.K

Alan Morris, B.A., D.Phil.

Department of Biological SciencesUniversity of Warwick

Coventry, U.K

Miguel Ortin, M.D., Ph.D., M.R.C Path.

St George’s HospitalLondon, U.K

K John Pasi, M.B., Ph.D., F.R.C.P., F.R.C.Path., F.R.C.P.C.H.

Department of HaematologyThe Royal London HospitalLondon, U.K

Geoffrey D Poole, M.Sc., F.I.B.M.S

National Blood ServiceBristol, U.K

R Martin Rowan (deceased), F.R.C.P.(G), F.R.C.P.(E), F.R.C.Path.

TOA Medical Electronics (Europe)and

University of GlasgowGlasgow, Scotland, U.K

N Keith Shinton, M.D., F.R.C.P., F.R.C.Path.

Clinical Sciences, Research InstituteWarwick Medical School

University of WarwickCoventry, U.K

Jerry Spivak, M.D.

Division of HaematologyJohns Hopkins University Medical SchoolBaltimore Maryland

3393_C000.fm Page vi Monday, November 19, 2007 1:58 PM

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List of Comprehensive Entries

ABO(H) blood groups 1

Acquired aplastic anemia 6

Acquired immunodeficiency syndrome 9

Acute myeloid leukemia 25

Adverse reaction to drugs 42

Allogeneic stem cell transplantation 51

Anemia 63

Arterial thrombosis 86

Autologous blood transfusion 99

Autologous stem cell transplantation 101

Blood components for transfusion 116

Blood donation 126

Bone marrow 133

Cell-adhesion molecules 151

Cell cycle 154

Cell signal transduction 160

Cell surface receptors 171

Chronic myelogenous leukemia 187

Coagulation-factor concentrates 193

Cobalamins 200

Cold agglutinin disease 206

Colloids for infusion 208

Colony forming units 211

Complement 215

Cytokines 230

Cytotoxic agents 237

Deoxyribonucleic acid 246

Disseminated intravascular coagulation 255

Drugs used for hematological disorders 263

Erythrocytosis 280

Essential thrombocythemia 286

Factor VIII 294

Fanconi anemia 302

Fetal/neonatal transfusion 307

Fibrinogen 310

Fibrinolysis 314

Flow cytometry 321

Folic acid 326

Gene therapy 343

Genomics 347

Graft-versus-host disease 357

Granulocytes 360

Hemapheresis 373

Hematology 377 3393_C000.fm Page vii Monday, November 19, 2007 1:58 PM

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Hematopoiesis 381

Hemoglobin 399

Hemoglobinopathies 409

Hemolytic anemias 416

Hemolytic disease of the newborn 420

Hemophilia A 427

Hemorrhagic disorders 432

Hemostasis 435

Heparin 438

Hereditary hemochromatosis 443

Histiocytes 451

Hodgkin disease 457

Human immunodeficiency viruses 463

Human leukocyte antigen 468

Immune thrombocytopenic purpura 486

Immunodeficiency 490

Immunoglobulins 496

Infectious mononucleosis 508

Inflammation 510

Interleukins 514

Ionizing radiation 526

Iron 527

Iron deficiency 531

Leukemogenesis 549

Lymphocytes 570

Lymphoproliferative disorders 580

Malaria 593

Megaloblastosis 607

Molecular genetic analysis 618

Monoclonal antibodies 622

Monoclonal gammopathies 624

Myelodysplasia 633

Myeloma 640

Myeloproliferative disorders 644

Neonatal hematology 651

Neutropenia 657

Neutrophil 663

Neutrophilia 669

Non-Hodgin lymphoma 673

Paroxysmal nocturnal hemoglobinuria 692

Plasmacytoma 706

Platelet 707

Platelet function disorders 711

Point-of-care-testing 721

Polycythemia rubra vera 723

Porphyrias 727

Pregnancy 734

Pretransfusion testing 738

Prion disease 742

Protein C 745

Pure red cell aplasia 755 3393_C000.fm Page viii Monday, November 19, 2007 1:58 PM

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Red blood cell 766

Red blood cell transfusion 777

Reticulocytes 790

Rhesus (Rh) blood groups 793

Sickle cell disorder 806

Spleen 819

Thalassemia 836

Thrombocytopenia 843

Thrombosis 849

Thrombotic thrombocytopenic purpura 851

Transfusion-transmitted infection 862

Vascular endothelium 877

Venous thromboembolic disease 883

Viral infection disorders 888

Von Willebrand disease 895

Warfarin 899

Warm autoimmune hemolytic anemia 904 3393_C000.fm Page ix Monday, November 19, 2007 1:58 PM

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List of Illustrations

1 Diagrammatic representation of ABH blood group biochemistry 2

2 Schematic diagram of proximal events in immune-mediated marrow failure 7

3 Changes in CD4 cell counts (●) and plasma viremia () during HIV infection 10

4 Morphological changes in apoptosis 84

5 Separation of whole blood into components 118

6 Simplified scheme of plasma fractionation 121

7 Diagrammatic representation of the structure of bone marrow at a diaphysis 134

8 The integrin receptor family 152

9 Phases of the cell cycle: restriction point 155

10 Phases of the cell cycle: early G1-S transition 156

11 Phases of the cell cycle: late G1-S transition 156

12 Phases of the cell cycle: G2-M transition 157

13 The stages of mitosis 158

14 Intracellular signal transduction pathways 161

15 Ras-mediated signal transduction pathways 164

16 Role of Janus kinases (JAKs) and signal transducers and activators of transcription (STATs) in cytokine signaling 167

17 The canonical Wnt signaling pathway 169

18 Cross talk between tumor cells, endothelial cells, and stromal cells is modulated by the Notch pathway and stimulates tumor angiogenesis 170

19 Proposed mechanisms of the Hh pathway activation 171

20 G-protein-coupled receptors Classical examples of seven-transmembrane (7TM)-receptor signaling 172

21 Cytokine receptor superfamily type 2 174

22 Role of checkpoint kinases in DNA damage 177

23 Structure of methylcobalamin 200

24 Bone marrow colony forming units (in vitro) 211

25 A schema of contact activation 221

26 Hypothetical genealogy of hematopoietic tissue-supporting cells 243

27 Structure of DNA 247

28 Replication of DNA 248

29 Domain structure of factor V and factor VIII 292

30 Model of factor VIII and thrombin cleavage sites 294

31 Central role of factor X in the final common pathway of coagulation 297

32 Activation of factor XI 299

33 Conversion of fibrinogen to cross-linked fibrin 311

34 Plasmin digestion of fibrinogen and non-cross-linked fibrin by plasmin 312

35 Plasmin digestion of cross-linked fibrin 313

36 Normal fibrinolysis 315

37 Conversion of plasminogen to plasmin 316

38 Diagrammatic representation of cell-counting and cell-volume measurement by aperture impedance and the effect of nonaxial flow 323

39 Schematic illustration of a hydrodynamic-focused (sheathed flow) counter using the light-scattering principle 323 3393_C000.fm Page x Monday, November 19, 2007 1:58 PM

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40 Diagrammatic representation of cytometry by light scattering 324

41 Structure of folic acid 327

42 Reduced forms of folic acid 327

43 Compounds of folic acid 328

44 Folates in metabolic pathways 328

45 Gene-expression analysis using oligonucleotide microarray 348

46 Gene-expression analysis in two tissue samples using spotted DNA microarray 349

47 Granulocyte compartments 360

48 Changes in nuclear shape during neutrophil maturation 361

49 The Latham bowl 374

50 Temporal appearance of different types of assayable hematopoietic cells 382

51 Migration of progenitors that leads to definitive hematopoiesis 383

52 Schematic view of transcription factors and hematopoiesis 383

53 Stages of hematopoiesis in the embryo and fetus 385

54 Proportions of the various human hemoglobin polypeptide chains through early life 385

55 Proposed hierarchy of hematopoietic colony-forming potential 389

56 Molecular regulation of hematopoietic stem-cell niche 392

57 Surface phenotype of mouse and human hematopoietic stem cells 395

58 Structure of heme 397

59 Chemical steps in the biosynthesis of hemoglobin 398

60 Diagrammatic representation of the tertiary configuration of the myoglobin of sperm whale 400

61 Diagrammatic representation of the relationship between α- and β-globin chains in the hemoglobin tetramer 400

62 Steps in the development of hemoglobin shown diagrammatically 402

63 Oxygenated and deoxygenated hemoglobin molecule 402

64 Oxygen-dissociation curve 403

65 Degradation of hemoglobin 404

66 Diagrammatic representation of the overall hemostatic response following vessel wall injury 435

67 Classical “waterfall” hypothesis of coagulation 436

68 A revised coagulation cascade 437

69 Human immune deficiency virus (HIV) life cycle 464

70 The evolution of plasma laboratory markers of the naturalization of HIV infection 466

71 Induction of an immune response 467

72 Purine metabolism to uric acid 478

73 Schematic diagram of an immunoglobulin 497

74 Schematic diagram of light- and heavy-chain genes in mature B-cells 500

75 The structure of the V-Cµ-Cδ region of the heavy-chain genes of the mature B-cell 500

76 An IgG antibody positioned between two RBCs at the minimum distance of approach 506

77 Degradation of sphingolipids 561

78 Diagrammatic representation of the structure of a liver lobule 564

79 Diagrammatic representation of the lymph-node structure 568

80 Development of T-cells 573

81 Development of B-cells 573

82 Development of NK-cells 574 3393_C000.fm Page xi Monday, November 19, 2007 1:58 PM

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83 Flow diagram of investigation of macrocytic anemias 592

84 Life cycle of malaria plasmodium 593

85 Flow diagram of the investigation of microcytic anemia 614

86 Cycle of mitotic nuclear division 616

88 Polymerase chain reaction 620

89 Types of serum monoclonal protein 625

90 Diagnoses in patients with monoclonal protein 625

91 Relationship of the origin of myeloproliferative disorders 645

92 Simplified schematic diagram of the pathophysiology of neutropenia 657

93 Mechanisms of neutrophilia 669

94 Flow diagram of the investigation of normocytic anemia 680

95 Diagrammatic representation of nuclear shapes in Pelger-Huët anomaly 697

96 Diagrammatic representation of platelet ultrastructure 708

97 Cell signal transduction in the platelet-release reaction 710

98 Examples of platelet aggregation tracings 716

99 Structural formula of the porphyrin nucleus and diagrammatic representation of some naturally occurring porphyrins 730

100 Schematic diagram of the prion hypothesis 742

101 Diagrammatic representation of protein C and protein S activity 746

102 The proteomic process 749

103 Comparison of MALDI and SELDI mass spectroscopy 750

104 Diagrammatic representation of prothrombin activation 751

105 Red blood cell membrane cytoskeleton proteins 767

106 Metabolism of glucose by red blood cells 770

107 Diagrammatic representation of the histological structure of the spleen 820

108 Hypothetical stem cell development 827

109 Chart of thromboelastogram tracing 847

110 Diagrammatic representation of the structure of the thymus 853

111 Transcriptional regulation 860

112 Translation initiation in eukaryotes 866 3393_C000.fm Page xii Monday, November 19, 2007 1:58 PM

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List of Tables

1 Frequencies of the Major ABO Blood Groups 3

2 Laboratory Determination of ABO Blood Groups 4

3 Drugs Associated with Acquired Aplastic Anemia 6

4 Definition of Disease Severity of Aplastic Anemia (AA) 8

5 Morphological Features of Acute Lymphoblastic Leukemia 18

6 Classification and Incidence of Acute Lymphoblastic Leukemias by Immunophenotyping 19

7 Chromosomal Translocations and Genetic Alterations in Acute Lymphoblastic Leukemia 19

8 Prognostic Factors in Acute Lymphoblastic Leukemia 23

9 Association of FAB Subtype with Phenotypic Expression in Acute Myeloid Leukemia 26

10 Association of FAB Type with Chromosomal Abnormalities in Acute Myeloid Leukemia 28

11 Laboratory Features in Patients with Disseminated Intravascular Coagulation (DIC) in Acute Promyelocytic Leukemia 30

12 Hematological Adverse Reactions to Drugs 42

13 Drugs Commonly Causing Agranulocytosis 47

14 Hematological Effects of Alcohol 49

15 Indications for Allogeneic Stem Cell Transplantation, by Disease 52

16 Morphologic Classification of Anemia 65

17 Common Causes of Anemia by Morphology 65

18 Causes of Aplastic Anemia 82

19 Risk Factors for Arterial Thrombosis 88

20 Classification of Autoimmune Hemolytic Anemias 98

21 Indications for Autologous Blood Transfusion Procedures 99

22 Comparison of Intraoperative Blood Salvage (IBS) Systems 101

23 Advantages and Disadvantages of Autologous and Allogeneic Stem Cell Transplantation 102

24 Conditioning Protocols for Autologous Stem Cell Transplantation Used in Acute Myeloid Leukemia 102

25 Granules of Basophils and Mast Cells 110

26 Causes of Basophilia 111

27 Composition of Platelet Concentrates 120

28 Patients at Risk for TA-GVHD 124

29 Properties of Some Less Commonly Encountered Antigens and Antibodies of Blood Group Systems 130

30 Cytochemical and Histochemical Stains and Their Identifications 138

31 Causes of Bone Marrow Hypoplasia 140

32 The β1 (CD29, very late antigen [VLA]) Integrin Family 152

33 The β2 (CD18, leukocyte integrins) Integrin Family 153

34 The β3 (CD61, cytoadhesin) Integrin Family 153

35 The Selectin Family 154

36 Cytokine Signaling Abnormalities 171 3393_C000.fm Page xiii Monday, November 19, 2007 1:58 PM

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37 Staging of Chronic Lymphocytic Leukemia 187

38 Diagnostic Criteria for Chronic Neutrophilic Leukemia 191

39 Coagulation Factors and Their Synonyms 198

40 Ligands of Cobalamin 200

41 Typical Compositions of Human Albumin Solutions 209

42 Cytokine Nomenclature: Early Interleukins 232

43 Cytokine Nomenclature: Hematopoietic Factors 232

44 Cytokine Nomenclature: Nonhematopoietic Growth Factors and Other Growth Regulatory Cytokines 232

45 Cytokine Nomenclature: Antiviral Cytokines — Interferons (IFNs) 233

46 Cytokine Nomenclature: Tumor Necrosis Factors 233

47 Cytokine Nomenclature: Cytokines Involved in Pregnancy and Proliferation of Embryonic Cells 233

48 Summary of Major Cytokine Functions 234

49 Cytotoxic Combination Therapy 238

50 Reference Ranges in Health of Differential Leukocyte Counts (× 109/l) 252

51 Hemostatic Abnormalities in DIC 257

52 Clinical Manifestations of DIC 258

53 Drugs Associated with Vascular Purpura 263

54 Commonly Used Drugs in Hematological Disorders 263

55 Phenotypes and Genotypes of the Duffy (Fy) Blood Group System 266

56 Causes of Dyserythropoiesis 267

57 Platelet Abnormalities in Dysmegakaryopoiesis 269

58 Content of Eosinophil Granules 274

59 Inflammatory Mediators 276

60 Erythrocyte Sedimentation Rate Ranges in Health 280

61 Causes of Erythrocytosis 281

62 Clinical Features of Fanconi Anemia 302

63 Properties of Fc Receptors 304

64 Clinical Disorders Related to Abnormal Fibrinolysis 319

65 Common Fluorochromes 324

66 Folates and Their Activities 327

67 A Grading System for Chronic Graft-versus-Host Disease 358

68 Causes of Granulomata Formation 363

69 Disorders Causing Heinz Body Formation 371

70 Adverse Effects of Apheresis Donation 374

71 Temporal Expression of Globin Chains 386

72 Hematopoietic Growth Factors 390

73 Hematopoietic Regulators 391

74 Structure of Physiological Hemoglobin Chains 401

75 Reference Ranges in Health of Hemoglobin in Peripheral Blood 404

76 Classification of Hemolytic Anemias 418

77 Hereditary Hematological Disorders 443

78 Specialized Organ Role of Histiocytes 454

79 Estimated Regional Incidence of HIV Infection by 2005 468

80 Action of Drugs on Immune Functions 494

81 Relationship of Immunodeficiency with Common Infections 495

82 Antibiotics Appropriate to Opportunistic Infections 496

83 IgG Subtype Properties 498

84 Causes of Ineffective Erythropoiesis 507 3393_C000.fm Page xiv Monday, November 19, 2007 1:58 PM

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85 Roles of the Tissue Macrophages (Histiocytes) in Inflammation 511

86 Commonly Used Therapeutic Interferons and Their Brand Names 514

87 Summary of Sources, Stimuli, and Biological Activity of Interleukins 515

88 Dose-Related Effects of Penetrating Ionizing Radiation 526

89 Common Phenotypes and Genotypes of the Kell Blood Group System 538

90 Phenotypes and Genotypes of the Kidd (Jk) Blood Group System 540

91 Summary of Hematological and Biochemical Changes Induced by Lead Toxicity 546

92 Common Karyotypic Abnormalities and Molecular/Functional Correlates 549

93 Causes of Leukemoid Reactions 555

94 Causes of Leukoerythroblastic Anemia 559

95 Phenotypes and Genotypes of the Lewis Blood Group System 560

96 Characterizing Markers of Mature Resting Lymphocytes 572

97 Genotypic and Cytogenetic Changes of Lymphoproliferative Disorders 582

98 REAL Classification of Lymphoid Tumors 584

99 Comparison of Commonly Used Lymphoma Classifications 585

100 WHO Classification of Tumors of Lymphoid Tissue 585

101 Predominant Sites of Selected Lymphoproliferative Disorders at Presentation 587

102 Ann Arbor Staging System of Lymphomas 587

103 Immunophenotypes of Lymphoproliferative Disorders 588

104 Febrile Illness Depending upon Type of Plasmodium 595

105 Microscopic Differentiation of Malaria Plasmodia 596

106 Hematological Changes Associated with Malignancy 598

107 Causes of Megaloblastosis 607

108 Causes of Microcytosis 614

109 Phenotypes and Genotypes of the MNS Blood Group System 617

110 Therapeutic Use of Monoclonal Antibodies 623

111 Causes of Monocytosis 629

112 Forms of Mucopolysaccharidosis 631

113 FAB Classification of Myelodysplasia 634

114 Targets of Interaction and Drugs under Trial in Treatment of Myelodysplasia 638

115 Causes of Secondary Myelofibrosis 639

116 Clinical and Laboratory Features of Myeloproliferative Disorders 647

117 Reference Ranges of Common Neonatal Blood Parameters 652

118 Causes of Neonatal Thrombocytopenia 654

119 Causes of Neutropenia 658

120 Antineutrophil Antibodies 661

121 Content of Neutrophil Granules 663

122 Killing Mechanisms of Neutrophils 665

123 Causes of Neutrophilia 670

124 Non-Hodgkin Lymphoma Grading According to the Working Formulation 674

125 Disorders Induced by Parvovirus B19 Infection 694

126 Phenotypes of the P and Associated Blood Group Systems 695

127 Important Platelet Membrane Glycoproteins 708

128 Disorders of Platelet Function 712

129 Guidelines for Transfusion of Platelets 719

130 Diagnostic Scoring System for Polycythemia Rubra Vera 725

131 Laboratory Features of the Porphyrias 727 3393_C000.fm Page xv Monday, November 19, 2007 1:58 PM

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132 Association of Enzyme Deficiency with Clinical Porphyria 728

133 Drug Groups Associated with Acute Porphyria 729

Individual Drugs Associated with Acute Porphyria 729

134 Protozoan Infections Causing Hematological Disorders 753

135 Causes of Pure Red Cell Aplasia 755

136 Major Red Blood Cell Membrane Proteins 768

137 Ideal Characteristics of Blood Substitutes 781

138 Available Hematology Reference Materials 783

139 Causes of Reticulin Fibrosis of Bone Marrow 790

140 Genotypes and Phenotypes of the Rh Blood Group System 793

141 Coagulation Changes Induced by Various Snake Venoms 817

142 Clinical Phenotypes and Genotypes of Thalassemias 837

143 α-Thalassemia Syndromes 838

144 Severity of β-Thalassemias 839

145 Genotypes of β-Thalassemia Syndromes 839

146 Causes of Thrombocytopenia 844

147 Drugs Associated with Thrombocytopenia 845

148 Causes of Thrombocytosis 846

149 Forms of ADAMTS 13 Deficiency 851

150 Microorganisms Transmissible by Blood Transfusion 862

151 Risk of Transmission of Certain Viruses 863

152 Classification of Vascular Purpura 880

153 Syndromes of Vasculitis 881

154 Ranges for Normal and Abnormal Levels of Viscosity 892

155 Classification of Von Willebrand Disease 895

156 Classification of Von Willebrand Disease Type 2 897

157 Management of Hemorrhage Due to Excess Warfarin 901

Reference Range Tables I Reference Ranges of Hematological Values for Adults 945

II Hemoglobin Concentrations (g/dl) for Iron-Sufficient Preterm Infants 946

III Hemoglobin F and Hemoglobin A2 in the First Year of Life (Measured by Electrophoresis) 947

IV Red Blood Cell Reference Ranges on First Postnatal Day during the Last 16 Weeks of Gestation 947

V Red Blood Cell Reference Ranges from Birth to 18 Years 947

VI Mean Serum Iron and Iron Saturation Percentage 948

VII Values of Serum Iron, Total Iron-Binding Capacity, and Transferrin Saturation from Infants during the First Year of Life 948

VIII Red Blood Cell Enzyme Activities 949

IX Peripheral Blood Leukocytes Reference Ranges (×109/l) 949

X Reference Ranges of Platelets in Peripheral Blood 950

XI Reference Ranges for Folic Acid (μg/l) 950

XII Bone Marrow Cell Populations of Normal Infants in Tibial Bone Marrow (mean % ± SD) 951 3393_C000.fm Page xvi Monday, November 19, 2007 1:58 PM

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XIII References Ranges for Coagulation Tests in Healthy Full-Term Infants

during First 6 Months of Life 952XIV Reference Ranges for Coagulation Tests in Healthy Premature Infants

(30–36 Weeks Gestation) during the First 6 Months of Life 952

XV Reference Ranges for Inhibitors of Coagulation in the Healthy Full-Term

Infant during the First 6 Months of Life (mean ± 1 SD) 953XVI Reference Ranges for Inhibitors of Coagulation in Healthy Premature

Infants during the First 6 Months of Life 9533393_C000.fm Page xvii Monday, November 19, 2007 1:58 PM

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ADCC antigen-dependent cellular cytotoxicity

AML acute myeloid leukemia; acute myeloblastic leukemia

ANCA antineutrophilic cytoplasmic antibodies

APC antigen-presenting cell; activated protein C

APRT adenine phosphoribosyltransferase (transaminase)

APSAC acylated plasma streptokinase activator complex

APTT activated partial thromboplastin time

ARAM antigen-recognizing activation motif

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AT/AT-III antithrombin-III

BCR break-point cluster region (genetics)

CDA congenital dyserythropoietic anemia; 2′-chlorodeoxyadenosine

CFU-GEMM colony forming unit-granulocytes/erythroid/macrophages/

megakaryocytesCFU-GM colony forming unit-granulocytes/macrophages

CFU-Meg colony-forming unit-megakaryocyte

3393_C000.fm Page xix Monday, November 19, 2007 1:58 PM

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CHAD cold hemolytic anemia disease

CHOP cyclophosphamide, adriamycin, vincristine, and prednisone or

CLEVER common lymphatic and endothelial vascular endothelial receptor

CML chronic myeloid leukemia; chronic myelogenous leukemia

CNSHA congenital nonspherocytic hemolytic anemia

CNTFR ciliary neurotrophic factor receptor

CSF colony stimulating factor; cerebro-spinal fluid

CSHH congenital self-healing histiocytosis

DILS diffuse infiltrative lymphocytosis syndrome

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DLBCL diffuse large B-cell lymphoma

EACA/EAPA epsilon-aminocaproic acid

EDRF endothelial-derived relaxing factor

FAB French-American-British (classification of leukemia)

FCAS familial cold-associated autoinflammatory disorder

FISH fluorescence in situ hybridization

FMAIT fetomaternal alloimmune thrombocytopenic purpura

rFVIII;c recombinant factor VIII coagulation

G-CSF granulocyte colony stimulating factor

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G-CSFR granulocyte colony stimulating factor receptor

GM-CSF granulocyte/macrophage colony stimulating factor

HAART highly active antiretroviral therapy

HAT/HIT heparin-associated/induced thrombocytopenia

acidified serum test

HGPRT hypoxanthine-guanine phosphoribosyltransferase

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HPA human platelet antigen

HPFH hereditary persistent fetal hemoglobin

HPP hereditary infantile pyropoikilocytosis

HPP-CFC high-proliferative-potential colony forming cell

HSCT hematopoietic stem cell transplantation

ICSH International Council for Standardization in Hematology

INR international normalized ratio (for prothrombin time)

IPSID immunoproliferative small intestinal disease

ISBT International Society of Blood Transfusion

ISTH International Society of Thrombosis and Hemostasis

3393_C000.fm Page xxiii Monday, November 19, 2007 1:58 PM

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IVT intravascular fetal transfusion

J-CML juvenile chronic myelocytic leukemia

LCAT lecithin cholesterol acyltransferase

LIFR leukocyte migration inhibiting factor receptor

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MRI magnetic resonance irradiation

MTP microsomal triglyceride transfer protein

NADPH nicotinamide adenine dinucleotide phosphate reduced

NAPTT nonactivated partial thromboplastin time

NHFTR nonhemolytic febrile transfusion reaction

NOD nucleotide-binding oligomerization domain protein

NRTi nonnucleoside reverse transcriptase inhibitor

PAS para-aminosalicylic acid; periodicacid-Schiff

PBSCT peripheral blood stem cell transplantation

PCI protein C inhibitor; percutaneous coronary artery intervention

PCR-SSCP polymerase chain reaction-single stranded conformational

polymorphisms

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PG prostaglandin (e.g., PGE1)

PIVKA protein-induced vitamin K absence (or antagonist)

POEMS polyneuropathy, organomegaly, endocrinopathy, monoclonal

gammopathy, and skin changes

PTTK partial thromboplastin time with kaolin

PUVA psoralen and ultraviolet-A radiation therapy

RAEB refractory anemia with excess blasts

rFVIII;c recombinant factor VIII coagulation

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RNI reactive nitrogen intermediate

SAG-M saline, adenine, glucose, and mannitol

S-phase cells in DNA-synthesis phase of cycle

STATS signal transducers and activators of transcription

TAFI thrombin-activatable fibrinolysis inhibitor

TA-GVHD transfusion-associated graft-versus-host disease

TAP transporters in antigen presentation; T-cell antigen presentation

TC-I, TC-II, TC-III transcobalamin I, II, III

TEC transient erythroblastopenia of childhood

Th/TH/TH T-lymphocyte helper cell

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TIL tumor-infiltrating lymphocytes

TRALI transfusion-related acute lung injury

TRAP tartrate-resistant acid phosphatase

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Preface ivNotice vContributors viList of Comprehensive Entries viiList of Illustrations xList of Tables xiiiAbbreviations xviiiA–Z 1APPENDIX I: WHO Classification of Tumors of Hematopoietic and

Lymphoid Tissues 911APPENDIX II: FAB Classification of Leukemia/Myelodysplasia 915Reference Range Tables 919References 929Internet References 951Bibliography 953Index 9553393_C000.fm Page xxix Monday, November 19, 2007 1:58 PM

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A monoclonal antibody that binds to glycoprotein IIb/IIIa (GpIIb/IIIa) on platelets andconsequently inhibits platelet function It is a Fab fragment of a human–mouse chimericmonoclonal antibody It has a very high affinity for (but a slow dissociation rate from) theGpIIb/IIIa Although it has a short half-life of 10 to 30 min, it has a long biological half-life due to its strong affinity to the GpIIb/IIIa receptor, which remains bound in circulationfor up to 15 days However, this prolonged presence has minimal residual activity, andplatelet function returns to baseline 12 to 36 h after therapy More than 80% of GpIIb/IIIamust be blocked to inhibit platelet function It has provided robust, consistent, and highlysignificant reductions in death or myocardial infarction in several large protein C inhibitor(PCI) trials; it is a preferred drug used during coronary interventions such as percutaneoustransluminal stenting Adverse drug reactions are those associated with hemorrhage and

hypersensitivity It should not be given to patients receiving drugs for activation of

fibrinolysis

ABETALIPOPROTEINEMIA

(Bassen-Kornweig syndrome) An autosomally recessive disorder caused by mutations of themicrosomal triglyceride transfer protein (MTP), resulting in defective synthesis of apoproteinB-100 in the liver and apoprotein B-48 in the interstitial cells Lipids, principally triglyceride,cholesterol, and cholesterol esters, are normally surrounded by a stabilizing coat of phospho-lipid Apoproteins are embedded in the surface of the complex of lipids and phospholipids

to form lipoproteins, which in this condition cannot be synthesized Low-density lipoproteins,such as chylomicrons, are absent from serum, leading to defective mobilization of triglycer-ides from the intestines and liver Plasma cholesterol and phospholipids are reduced, and theenzyme lecithin cholesterol acyltransferase (LCAT) is greatly reduced Triglycerides accumu-late in the absorptive cells of the small intestine, resulting in malabsorption in infancy withfailure to develop Fat intolerance and fat-soluble vitamin deficiency result in progressivecerebellar ataxia with peripheral neuropathy and retinitis pigmentosa Acanthocytes are seen

in the peripheral blood, their shortened red blood cell survival giving rise to a mild chronic

hemolytic anemia No specific treatment is available

ABO(H) BLOOD GROUPS Blood groups arising from two specific antigen–antibody systems located on red blood cells (RBCs) and platelets.

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2 ABO(H) BLOOD GROUPS

Biochemistry

The ABH antigens are carbohydrates, the A and B determinants being found at the termini

of the oligosaccharide chains of glycoproteins and glycolipids

Most ABH antigens on RBCs are glycoproteins, the carbohydrate portion usually beinglinked to protein, e.g., the anion transport protein (AE1 or Band 3) Because the ABH bloodgroup antigens are not localized to a particular protein, the antigens are also found widelydistributed in body tissues and occur in soluble form in secretions The presence or absence

of ABH substances in saliva determines the “secretor” status of the person (see Lewis blood groups)

Genetics and Phenotypes

The ABH antigens of RBCs are determined by genes belonging to the ABO and Hh bloodgroup systems Blood group genes in these systems specify glycosyl transferases thatcatalyze the transfer of monosaccharides from nucleotide sugars onto carbohydrate pre-cursors The product of the H gene is an alpha 1,2-L-fucosyl transferase The H antigenthat is produced is the biosynthetic precursor of the A and B antigens: the A gene specifies

an alpha 1,3-N-acetyl-D-galactosaminyltransferase, and the B gene specifies an D-galactosyltransferase (see Figure 1)

(alpha)1,3-FIGURE 1

Diagrammatic representation of ABH blood group biochemistry.

GalNAc Gal Carbohydrate Lipid or protein

Fuc L–fucose

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ABO(H) BLOOD GROUPS 3

The O gene is a common allele at the ABO locus on chromosome 9 and does not result

in the production of A or B antigens A number of alleles at the ABO locus result in aweakened or altered expression of A and B antigens For example, RBCs of the A3 and

Ax phenotypes react with some anti-A antibodies, but not with others Weak phenotypes

may also have a nongenetic cause, e.g., old age and disease (particularly leukemia) Table 1shows the incidence of the major ABO phenotypes These frequencies vary significantlybetween different ethnic groups; for example, the incidence of blood group B is muchhigher in Africans and in Vietnamese ABO polymorphisms have been claimed to haveputative relationships with disease.1

A very rare allele at the H locus on chromosome 19 is the h gene, which results in noexpression of H, and since H is the precursor for A and B, people with the genotype hhhave no A, B, or H antigens on their RBCs These people are said to have the “Bombay”(Oh) null phenotype, because of its relatively high frequency in Bombay, India

Antibodies and Their Clinical Significance

Alloantibodies to the A, B, and H antigens are naturally occurring They are found in theplasma of people who lack the corresponding antigen and who have had no known RBCstimulus, e.g., transfusion or pregnancy (see Table 1) Exposure to widely distributedbacteria carrying ABH-determinant structures accounts for the production of anti-A andanti-B in infants Maternal ABO antibodies are often found in the plasma of newborns.Anti-A and anti-B (and anti-H found in Bombay persons) are often active at 37°C andbind complement, and may give rise to severe hemolytic transfusion reactions if incom-patible RBCs are transfused (see Red blood transfusion) ABO antibodies may cause

hemolytic disease of the newborn, but this is rarely severe, because:

ABO antigens are not fully developed on fetal RBCs

ABO antigens are present on tissues other than RBCs, allowing neutralization of theantibody to occur

Maternal ABO antibodies may be partly or wholly IgM

Weak examples of anti-A, anti-H, and, occasionally, anti-B are sometimes found in theplasma of persons who have weak or altered expressions of the relevant antigens Forexample, anti-A1 may be found in people with the phenotypes A2B and Ax These anti-bodies are rarely of clinical significance, but may cause problems in pretransfusion testing

ABO Blood Grouping

Determination of ABO groups is an essential requirement of pretransfusion testing

Anti-A and anti-B reagents are used to define the antigens on the patient’s RBCs, and the

TABLE 1

Frequencies of the Major ABO Blood Groups

Phenotype Genotype

Frequency in Caucasians (%)

ABO Antibody Regularly Present in Plasma

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(Spur cells) Abnormal red blood cells (RBCs) characterized by having 2 to 20 irregularlyplaced spicules or thornlike projections of unequal length These spicules are caused bychanges in the lipid content of the red blood cell membrane, possibly from reduction ofglycophospholipids with relative increase of sphingomyelin Once produced, the change

is usually irreversible Acanthocytes are associated with other disorders in some patients.Hereditary

• Abetalipoproteinemia

• Surface antigen polymorphisms:

McLeod phenotype of the Kell blood group – X-linked anomalyInhibition of Lutheran blood groups Lua and Lubsomatic dominantanomaly

• Red blood cell membrane disorders

High RBC membrane phosphatidylcholineAbnormal RBC membrane Band 3 protein

Laboratory Determination of ABO Blood Groups

Patient’s Blood Group

anti-A + – – + Cell or forward group

(reagent antibody tested against patient’s RBCs)

A cells – + + – Serum or reverse group

(patient’s serum or plasma tested against reagent RBCs)

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ACIDIFIED SERUM LYSIS TEST 5

Acquired

Liver disorders, particularly alcoholic cirrhosis

• Malnutrition

HypobetalipoproteinemiaAnorexia nervosa, cystic fibrosis

• Hypothyroidism and panhypopituitarism

Infantile pyknocytosis

Vitamin E deficiency in premature newborns

Splenic hypofunction and splenectomy

Acanthocytes have a reduced red blood cell survival This may give rise to a mild chronic

hemolytic anemia (spur cell hemolytic anemia, Zieve’s syndrome)

ACETYLCHOLINESTERASE

(AChE) An enzyme of the erythrocyte (red blood cell) membrane that has greater activity

in young erythrocytes and decreases progressively with age This decrease in membraneAChE activity is a consistent erythrocyte abnormality in paroxysmal nocturnal hemoglo- binuria (PNH), reflecting defects in the phosphotidyl-inositol anchor

ACIDIFIED GLYCEROL LYSIS TEST

A screening test for hereditary spherocytosis In symptomless relatives of known cases,

it has a higher detection rate than the conventional osmotic fragility test of red blood cells, although the same principles apply, but it is also positive in other causes of sphero-cytosis It requires less blood than the osmotic fragility test and can be performed morerapidly Measurement is carried out in a recording spectrophotometer with the wavelengthset at 625 nm

ACIDIFIED SERUM LYSIS TEST

(Ham test) A test used for the diagnosis of paroxysmal nocturnal hemoglobinuria (PNH).2

Red blood cells from defibrinated, oxalated, citrated, or ethylenediamine tetraacetate(EDTA) anticoagulated patient’s blood are treated at 37°C with normal or patient’s serumacidified to pH 6.5 to 7.0 Normal serum known to be strongly lytic to PNH cells should

be used, but the test should be repeated with the patient’s own serum to exclude a form

of congenital dyserythropoietic anemia — HEMPAS (hereditary erythroblastic clearity associated with a positive acidified serum test) — in which lysis occurs in onlyabout 30% of cells with normal serum and not at all with the patient’s own serum.HEMPAS patients have a negative sucrose lysis test, but this test is positive in those withPNH The lytic potency of normal serum varies and is destroyed by mild heat, even atlow temperature within a few days, so the serum should be used fresh Lysis is measured

multinu-as liberated hemoglobin in a spectrophotometer at the wavelength of 540 nm In PNH, 10

to 15% lysis is usually obtained, with a range of 5 to 80% If the patient has been recentlytransfused, the degree of hemolysis will be reduced Confirmation of the diagnosis ofPNH requires the demonstration of GPI-linked molecules within the red blood cell mem-brane by flow cytometry

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6 ACQUIRED APLASTIC ANEMIA

ACQUIRED APLASTIC ANEMIA

(Idiopathic aplastic anemia) An uncommon disorder, with an incidence of 1 to 2 per millionper year in Western countries and perhaps twice or three times as common in China,Southeast Asia, and Japan.3 All ages are affected, with a peak incidence in young adultsand again in older patients In children and young adults, acquired aplastic anemia must

be distinguished from congenital types; in the elderly, it must be distinguished frommyelodysplastic syndrome The majority of cases (70%) are idiopathic; 15% follow expo-sure to drugs, which only rarely cause blood dyscrasias, and about 10% follow a hepaticillness, presumed to be viral, although no specific agent has yet been identified Theremaining cases follow other known viral infections or exposure to industrial or domesticagents Drugs associated with aplastic anemia are shown in Table 3

Pathogenesis

The disease is the result of dysfunction in the hematopoietic stem cell population There

is reduction in the number of CD34+ hematopoietic precursors Remaining cells have aproliferative defect The most convincing evidence suggests that the damage is caused byautoimmune cytotoxic T-cell attack In vitro evidence to support the autoimmune hypoth-esis has been summarized3–5a:

60 to 80% respond to immunosuppressive therapy using ATG and ciclosporin

Increased levels of cytokines that inhibit hematopoiesis (IFN-γ, TNF-α)

Increased Fas-antigen expression on bone marrow CD34+ cells

Activated cytotoxic CD8+ T-cells present in blood and bone marrow

quinacrine Nonsteroidal anti-inflammatory drugs phenylbutazone

indomethacin naproxen diclofenac ibuprofen piroxicam Antirheumatic gold salts

D-penicillamine

methylthiouracil propylthiouracil Psychotropic/antidepressants phenothiazines

mianserin dothiepin Anticonvulsants carbamazepine

phenytoin Antidiabetics chlorpropamide Carbonic anhydrase inhibitors acetazolamide

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ACQUIRED APLASTIC ANEMIA 7

Oligoclonal expansion of CD8+ T-cells

Upregulation of apoptosis and immune response genes

Human leukocyte antigen (HLA) restriction with overrepresentation of HLA-DR15

In acquired aplastic anemia, activated cytotoxic T-cells secrete cytokines such as tumornecrosis factor (TNF)-α and interferon (IFN)-γ, which inhibit hematopoietic progenitorcells (HPC) TNF-α and IFN-γ upregulate the expression of the Fas receptor on HPCs,triggering apoptosis Increased production of interleukin-2 results in expansion of T-cells.TNF-α and IFN-γ also increase nitric oxide (NO) production by marrow cells, which maycontribute to immune-mediated cytotoxicity and elimination of HPCs This is summarized

in Figure 2

Laboratory Features

The laboratory diagnosis depends upon the peripheral-blood cell levels, film examination, and bone marrow aspirate and trephine Other conditions that lead topancytopenia, particularly acute leukemias, must be excluded In the peripheral bloodthere is anemia, granulocytopenia, and a low platelet count Lymphocytes are normal innumber or reduced Typically there is some red blood cell macrocytosis and the reticulocytecount is low, either absolutely or in proportion to the degree of anemia Fetal hemoglobinlevels may be normal or slightly elevated, but these are not helpful in the differentialdiagnosis There may be some variation in size and shape of red cells, although this is notusually marked The presence of nucleated red cells raises the possibility of paroxysmalnocturnal hemoglobinuria (PNH) or myelodysplastic syndrome (MDS) Neutrophils arereduced, but their morphology is normal apart from some increase in granulation, so-called toxic granulation There is typically no shift to the left Platelet numbers are reduced,and the platelets are mostly small in size Abnormal cells including blasts or hairy cellsare significantly absent The diagnosis ultimately rests on bone marrow examination.Aspiration in typical cases is “easy.” The marrow films contain many fragments that are

peripheral-blood-of hypocellular, “lacy” appearance, and the cell trails are hypocellular, with mainly phocytes and nonhematopoietic cells remaining The bone marrow trephine is the keyinvestigation There is usually overall hypocellularity, with hematopoietic marrow

lym-FIGURE 2

Schematic diagram of proximal events in immune-mediated marrow failure: IFN- γ = interferon- γ ; IRF-1 = IFN regulatory factor-1; NOS = nitric oxide synthase; TNF = tumor necrosis factor (From Young, N.S., Rev Clin Exp Haematol., 4, 426–459, 2000 With permission.)

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8 ACQUIRED APLASTIC ANEMIA

replaced by fat cells, although cellular, or even hypercellular islands (“hot spots”), may

be found Reticulin is not increased, and no abnormal cells are seen Cytogenetic analysis

of the marrow is difficult, but it is usually normal, although abnormal clones, particularly

of trisomy 8 or monosomy 7, are occasionally found The significance is uncertain,

some-times indicating progression to MDS or acute leukemia There is no increased

chromo-somal instability in response to clastogens, as is seen in Fanconi anemia Mutations in

telomeres, with alterations in the ribonucleic protein complex responsible for synthesis

and maintenance of telomere repeats, has been reported.6

Clinical Features

The pancytopenia develops slowly, such that symptoms, which are the consequence of

the cytopenias, may be delayed.7 Purpura, easy bruising, or repeated epistaxis that is

difficult to control are common presentations Persistent sore throat with or without fever

and pallor, tiredness, or other effects of anemia may also be presenting features The course

of the untreated disease depends upon the severity of the bone marrow failure, as reflected

in the peripheral blood The degree of hypoplasia may remain stable over the ensuing

months or years or become more severe, necessitating therapeutic intervention

Paroxys-mal nocturnal hemoglobinuria and myelodysplasia, which may further progress to acute

myeloid leukemia, develop in 10 to 20% of patients Spontaneous recovery rarely occurs

Treatment

General Supportive Measures

Before specific treatment became available, only about 10% of patients with SAA or VSAA

(see Table 4) survived more than 1 year Current management depends on red blood cell

transfusion and platelet transfusion to correct the cytopenias and on prevention and

treatment of infection consequent upon any neutropenia. Definitive treatments to restore

stem cell function rely either on immunosuppression to allow partial or complete recovery

of peripheral blood pancytopenia, or stem cell transplantation to replace stem cells.8

Immunosuppression (IS)

The standard therapy is with intravenous immunoglobulin, antilymphocyte globulin

(ALG), given daily for 5 days (Some preparations are labeled antithymocyte globulin, or

ATG.) Ciclosporin, started after the ALG, may increase the rate and incidence of remission,

and some remissions are ciclosporin dependent About 65% of patients will achieve

remis-sion as defined as freedom from transfuremis-sion dependence Response is similar at all ages,

TABLE 4

Definition of Disease Severity of Aplastic Anemia (AA)

Severe (SAA) bone-marrow cellularity <25%, or 25–50% with <30% residual hematopoietic cells;

with 2/3 of the following:

neutrophils <0.5 × 10 9 /l platelets <20 × 10 9 /l reticulocytes <20 × 10 9 /l Very severe (VSAA) as for severe AA but with neutrophils <0.2 × 10 9 /l

Nonsevere (NSAA) patients not fulfilling the criteria for severe or very severe AA; with a hypocellular

marrow with 2/3 of the following:

neutrophils <1.5 × 10 9 /l platelets <100 × 10 9 /l hemoglobin <10 g/dl

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ACQUIRED IMMUNODEFICIENCY SYNDROME 9

and IS remains the treatment preferred for patients over the age of 40 years with severe

or very severe aplastic anemia, even if a suitable donor is available Improvement is slow,counts rarely improving before 6 weeks following ALG and may be delayed for 6 months

or more Second or third courses of IS may be effective in inducing remission in a fewpatients Relapse, defined as a return to transfusion dependence, occurs in up to 40% ofresponders over 10 years

High-dose cyclophosphamide, with or without ATG, apart from its use as a conditioningtherapy prior to transplantation, can be particularly useful when a suitable donor is

not available Monoclonal antibody therapy with anti-CD20 (rituximab) and anti-CD52 (alemtuximab) is being evaluated for refractory cases.

Allogeneic Hematopoietic Stem Cell Transplantation

(HSCT) See also Allogeneic stem cell transplantation; Hematopoietic stem cell assays When a human leukocyte antigen (HLA)-compatible sibling donor is available, HSCT is

the treatment of choice for children with SAA or VSAA and for adults under 40 years ofage with VSAA HSCT from unrelated, HLA-matched volunteer donors has become moresuccessful with the development of less-toxic conditioning regimens and is offered fol-lowing failure of IS treatment for younger patients Event-free survival is about 70% and

is age dependent, as with all HSCT Survivors do not have a higher incidence of abnormalclones of PNH or MDS cells arising Children have normal growth and fertility so long

as irradiation is avoided in conditioning for transplant

ACQUIRED HEMOLYTIC ANEMIA

See Hemolytic anemias.

ACQUIRED IMMUNODEFICIENCY SYNDROME

(AIDS) A clinical disorder arising from immunodeficiency following infection, usually by the human immunodeficiency virus-1 (HIV-1), less commonly by HIV-2, and rarely by

an undetermined cause The first patients to be infected were identified in 1981 Sincethen, the incidence has rapidly increased throughout the world; by 2003, an estimated

20 million had died from the disease Early diagnosis by HIV screening is recommended.20

Clinical Features

These occur in stages over a number of years Due to differences in immunologicalresponse, they must be considered separately for adults and infants

Adults

The best indicator of progression is the level of CD4 lymphocytes, as measured by flow

cytometry (see Figure 3)

Acute Seroconversion Syndrome

During the prodromal stage (10 to 40 days), evidence of viremia can be detected At theend of this time, approximately 50% of individuals develop an inflammatory illness similar

to infectious mononucleosis with transient lymphadenopathy, fever, pharyngitis, myalgia,

arthralgia, maculopapular skin eruptions, aphthous ulcerations, diarrhea, nausea, vomiting,and headache Neuropathic symptoms such as peripheral neuropathy or asymptomatic

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