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Uses of Beryllium As a result of beryllium’s unusual physical properties, such as its high melting point, high electrical conduc-tivity, high heat capacity, and oxidation resistance, be-

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percent for a total of about $10.7 million This

indus-try began in Belgium in 1807 when the British started

a blockade of cane sugar from the Caribbean during

the Napoleonic Wars With cane sugar unavailable,

beet sugar began to be the sugar of choice throughout

Napoleonic Europe The sugar production capital of

Belgium is Tienen, which hosts a large sugar-beet

pro-cessing factory that was founded in 1836 This factory

and related sugar production facilities owned by the

Raffinerie Tirlemontoise Group employ nearly two

thousand people This company owns three other

Belgian sugar factories, in Brugelette, Genappe, and

Wanze

Beer

Monks in Belgium began brewing beer sometime

dur-ing the Middle Ages There are more than one

hun-dred breweries scattered throughout Belgium, with

about eight hundred standard types of beer produced

These range from light through dark types of beer;

Belgians brew and export nearly every type of beer

possible Often, each type of beer is served in its own

distinctive glass, which is said to enhance the flavor

of that particular type of beer Though Belgium is

famous for many kinds of beer, it is possibly most

fa-mous for lambic beer, which is made in an ancient

brewing style This style depends on a spontaneous

natural fermentation process after ingredients are

ex-posed to the wild yeasts and bacteria native to the

Senne Valley, located south of Brussels This unusual

fermentation process produces a drink that is

natu-rally effervescent or sparkling, which is then aged, up

to two or three years, to improve its taste Much like

champagne (only produced in a certain region in

France) or Madeira (only produced on a certain

is-land owned by Portugal), the title of “lambic beer”

can only be given to this type of beer brewed in the

small Pajottenland region of Belgium Nearly half of

the beer brewed in Belgium is exported, mostly to

Canada, France, Germany, Italy, Spain, the United

States, and the United Kingdom

Chocolate

During the seventeenth century, when the Low

Coun-tries were ruled by Spain, Spanish conquistadores

brought cacao beans back from the New World to the

region that is now Belgium By 1840, the Berwaerts

Company had begun to sell Belgian chocolates that

were quite popular However, not until the nineteenth

century, when King Leopold II colonized the Belgian

Congo in 1885 and discovered cacao tree fields there, did Belgian chocolatiers begin to manufacture Bel-gian chocolates on a large scale At the beginning of the 1900’s, there were at least fifty chocolate makers in Belgium In 1912, Jean Neuhaus created a process for making a chocolate shell that could be filled with any number of fillings, something he called a “praline,” making Belgian chocolates even more popular Bel-gium produces more than 156,000 metric tons of chocolate each year, has more than two thousand chocolate shops throughout the country, and hosts about three hundred different chocolate companies Many of the original chocolate-making companies— such as Godiva, Leonidas, Neuhaus, and Nirvana— are still in operation today, and many of them still make chocolates by hand, using original equipment, high-quality ingredients, and Old World manufactur-ing techniques Chocolate shops in Belgium offer tast-ings, much like wineries, and host chocolate festivals, workshops, tours, and demonstrations There is a mu-seum dedicated to chocolate, the Musée du Cacao et

du Chocolat, near the Grand Place, the town square

in Brussels Belgium’s European Union neighbors (particularly France, Germany, and the United King-dom) are the biggest importers of Belgian chocolate

Pharmaceuticals Belgium has become a world leader in the pharma-ceutical industry, employing nearly thirty thousand people and accounting for about 10 percent of all Bel-gian exports Major pharmaceutical companies head-quartered in Belgium include UCB, Solvay, Janssen Pharmaceutica, Omega Pharma, Oystershell NV, and Recherche et Industries Thérapeutiques Private in-vestment in research and development in the phar-maceutical industry is at about 40 percent, which is nearly twice the average of other European compa-nies The pharmaceutical industry is also heavily sup-ported by the Belgian government, which offers tax incentives for pharmaceutical research and develop-ment The United States has imported about $2.3 bil-lion annually in medicinal, dental, and pharmaceuti-cal products from Belgium, which accounts for about

16 percent of all exports from Belgium to the United States

Textiles Since the thirteenth century, Belgium has been known as the home of master textile producers The famous Unicorn Tapestries or “The Hunt of the

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corn” series on display at The Cloisters, a part of the

Metropolitan Museum of Art in New York, is thought

to have been woven in Brussels sometime around

1495-1505 (when that area was still part of the

Nether-lands) The Flanders, or Flemish, region of Belgium is

still home to many lace-making artists, particularly in

the area of Bruges, which is the home of bobbin lace;

however, lace is also still produced in Brussels and

Mechelen This industry can be traced back to the

fif-teenth century, when Charles V decreed that lace

making was to be taught in the schools and convents

of the Belgian provinces to provide girls with a source

of income, as lace was popular on collars and cuffs for

clothing of both sexes at that time Lace is still

pro-duced in Belgium by lace artisans in their homes, one

piece at a time, and, thus, is a source of artistic lace

rather than high-production lace There is even a

mu-seum dedicated solely to lace, the Musée du Costume

et de la Dentelle, located near the Grand Place Other

textile production, including cotton, linen, wool, and

synthetic fibers, is concentrated in Ghent, Kortrijk,

Tournai, and Verviers, where carpets and blankets are

manufactured

Other Resources

As mentioned above, Belgium has few natural

re-sources, and its economy depends on importing raw

materials, processing those materials, manufacturing,

and exporting a finished product However, in

addi-tion to sugar processing, there are a few agricultural

resources grown and exported by Belgian farmers

These include fruits, vegetables, grains (wheat, oats,

rye, barley, and flax), tobacco, beef, veal, pork, and

milk

Other industries in which Belgian workers are

in-volved in processing imported goods that are then

ex-ported are motor vehicles and other metal products,

scientific instruments, chemicals (fertilizers, dyes,

plas-tics), glass, petroleum, textiles, electronics, and

pro-cessed foods and beverages, such as the beer and

chocolate described above

Marianne M Madsen

Further Reading

Binneweg, Herbert Antwerp, the Diamond Capital of the

World Antwerp: Federation of Belgian Diamond

Bourses, 1993

Blom, J H C., and Emiel Lamberts History of the Low

Countries New York: Berghahn Books, 2006.

Hieronymus, Stan Brew Like a Monk: Trappist, Abbey,

and Strong Belgian Ales and How to Brew Them

Boul-der, Colo.: Brewers, 2005

Kockelbergh, Iris, Eddy Vleeschdrager, and Jan

Wal-grave The Brilliant Story of Antwerp Diamonds

Ant-werp: MIM, 1992

Mommen, Andre The Belgian Economy in the Twentieth Century New York: Routledge, 1994.

Parker, Philip M The 2007 Import and Export Market for Unagglomerated Bituminous Coal in Belgium San

Diego, Calif.: ICON Group International, 2006

Sparrow, Jeff Wild Brews: Culture and Craftsmanship in the Belgian Tradition Boulder, Colo.: Brewers, 2005 Wingfield, George Belgium Edgemont, Pa.: Chelsea

House, 2008

Witte, Els, Jan Craeybeckx, and Alain Maynen Politi-cal History of Belgium: From 1830 Onwards Brussels:

Free University of Brussels Press, 2008

Web Sites Belgium: A Federal State http://www.diplomatie.be/en/belgium U.S Department of State

Background Note: Belgium http://www.state.gov/r/pa/ei/bgn/2874.htm See also: Coal; Diamond; Sugars; Textiles and fab-rics

Beryllium

Category: Mineral and other nonliving resources

Where Found The element beryllium is believed to occur in the Earth’s igneous rocks to the extent of 0.0006 per-cent It does not occur in its free state in nature; it is found only in minerals The leading producers are the United States, China, and some African countries

Primary Uses Beryllium has a number of important industrial and structural applications Its widest use is in the prep-aration of alloys used in the manufacture of watch springs, welding electrodes, hypodermic needles, den-tures, and molds for casting plastics Metallic beryl-lium is used to make windows in X-ray tubes because

of its high degree of transparency Finally, beryllium

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compounds have various uses in glass manufacture, in

aircraft spark plugs, and as ultra-high-frequency radar

insulators

Technical Definition

Beryllium (abbreviated Be), atomic number 4,

be-longs to Group II of the periodic table of the elements

and is one of the rarest and lightest structural metals

It has four naturally occurring isotopes and an

aver-age atomic weight of 9.0122

Description, Distribution, and Forms

Pure beryllium is a steel-gray, light, hard, and brittle

metal that becomes ductile at higher temperatures

and may be rolled into a sheet Beryllium burns with a

brilliant flame, but it becomes oxidized easily and

forms a protective coating of the oxide Beryllium has

a density of 1.85 grams per cubic centimeter, a

melt-ing point of 1,285° Celsius, and a boilmelt-ing point of

2,970° Celsius

Among the elements, ber yllium ranks

thirty-second in order of abundance Like lithium, it is

usu-ally isolated from silicate minerals It is believed that

its nucleus, like the nucleus of lithium and boron, is

destroyed by high-energy protons in the Sun and

other stars As a result it cannot survive the hot, dense

interiors of the stars, where elements are formed,

which accounts for its low abundance At least fifty

beryllium-containing minerals are known, but only

beryl and bertrandite—which contain up to 15

per-cent beryllium oxide and whose clear varieties are the

gems aquamarine and emerald—are the major

pro-ducers of the metal The richest beryllium-containing

ore deposits are pegmatite varieties of granite rocks

Many beryllium compounds have properties that

re-semble those of aluminum compounds Beryllium

ox-ide absorbs carbon dioxox-ide readily and is moisture

sensitive Beryllium hydroxide is a gelatinous

precipi-tate that is easily soluble in acid All beryllium halides

are easily hydrolyzed by water and emit hydrogen

halides

History

Beryllium was discovered as an oxide by Louis-Nicolas

Vauquelin during an analysis of emerald in 1798 and

was originally named glucinum because of the sweet

taste of its salts It was first isolated as a free metal by

Friedrich Wöhler and Antoine Bussy, who reduced

be-ryllium chloride with potassium metal

Obtaining Beryllium Beryllium ore is usually converted to a more reactive compound, such as beryllium fluoride, which is then electrolyzed with magnesium The element is inert with respect to water

Beryllium exists in the atmosphere of urban and coal-burning neighborhoods in much greater quanti-ties than in rural areas Dry dust, fumes, and aqueous solutions of the metal compounds are toxic, creating dermatitis, and inhaling them produces the effects of phosgene gas Its toxicity is believed to result from the substitution of the smaller beryllium atoms for mag-nesium atoms in enzymes, which are the biochemical catalysts

Uses of Beryllium

As a result of beryllium’s unusual physical properties, such as its high melting point, high electrical conduc-tivity, high heat capacity, and oxidation resistance, be-ryllium serves as a component in alloys of elements such as copper, where it adds a high tensile strength to

Aerospace 10%

Electrical components 22.5%

Electronic components 62.5%

Other 5%

Source:

Historical Statistics for Mineral and Material Commodities in the United States

U.S Geological Survey, 2005, beryllium statistics, in

T D Kelly and G R Matos, comps.,

, U.S Geological Survey Data Series 140 Available online at http://pubs.usgs.gov/ds/2005/140/.

U.S End Uses of Beryllium

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the metal The added beryllium is no more than 3

per-cent of the alloy Beryllium’s ability to transmit X rays

seventeen times more effectively than aluminum

makes it useful in cases where high-intensity X-ray

beams are needed

Soraya Ghayourmanesh

Web Sites

U.S Department of Labor: Occupational

Safety and Health Administration

Safety and Health Topics: Beryllium

http://www.osha.gov/SLTC/beryllium/

U.S Geological Survey

Minerals Information: Beryllium Statistics and

Information

http://minerals.usgs.gov/minerals/pubs/

commodity/beryllium/

See also: Alloys; Boron; China; Lithium; Nuclear

en-ergy; United States

Bessemer process

Category: Obtaining and using resources

The Bessemer process was the first method for produc-ing large quantities of inexpensive steel.

Definition

In the 1850’s, Henry Bessemer, looking for a way to improve cast iron, stumbled upon a way to make a new kind of steel By blowing air through molten iron in a crucible, he was able to burn off the carbon and many harmful impurities, and then the iron was heated to the point that it could be poured into molds Bessemer eventually learned to add Spiegeleisen, a manganese-rich cast iron, to the molten iron after the carbon and impurities were burned off The manga-nese countered the effects of the remaining traces of oxygen and sulfur, while the carbon (always present in cast iron) helped create the properties of steel

The Bessemer converter, on display at England’s Science Museum, was used for steel production and is recognized as an important invention

of the Industrial Revolution (SSPL via Getty Images)

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Prior to the late 1850’s, there were two common

iron-based construction materials One was cast iron, an

impure, brittle, high-carbon material used in

col-umns, piers, and other load-bearing members The

other was wrought iron, a workable, low-carbon

mate-rial used in girders, rails, and other spans The word

“steel” usually referred to a custom material produced

in very small quantities by adding carbon to

high-quality wrought iron

Bessemer’s resulting product, which came to be

known as “mild steel,” proved to be reliable and

dura-ble Because of these qualities, and because it could

be produced in large quantities, mild steel quickly

found widespread use in rails, ship plates, girders, and

many other applications, often replacing wrought

iron

Brian J Nichelson

See also: Iron; Manganese; Metals and metallurgy;

Steel

Biodiversity

Category: Ecological resources

Scientist Walter G Rosen coined the term

“biodiver-sity” in 1986 for the National Forum on Biodiversity;

the term was popularized later by the biologist Edward

O Wilson Biodiversity includes the variations and

associated processes within and among organisms It

is linked to the stability and predictability of ecosystems

and can be measured through the numbers and

compo-sition of species.

Background

Conservation was a priority in the United States in the

late 1800’s and early 1900’s, but efforts were driven by

the mistaken beliefs that there were regions

un-touched by humanity and that humans were not part

of nature Intensified use of lands leading up to and

during World War II hastened the loss of species and

wilderness areas The science of ecology was

emerg-ing but “natural” ecosystems were hard to identify

Thus, conservation efforts in the 1960’s and 1970’s

fo-cused on the preservation of particular species in

or-der to preserve biodiversity and led to passage of the

Endangered Species Preservation Act in 1966

Politi-cal support for protecting the environment and biodi-versity spread globally, leading to the 1992 Earth Sum-mit, in which representatives of 175 nations met in Rio

de Janeiro, Brazil As of 2009, all countries present at the summit, except the United States, had ratified the agreements All participating countries were expected

to identify, monitor, and report on various aspects of biodiversity within their borders; help deteriorating regions recover; include indigenous peoples in dis-cussions of biodiversity; and educate citizens about the importance of biodiversity Preservation of origi-nal habitats was preferred over off-site recovery ef-forts

Recognizing and Measuring Biodiversity Biodiversity can be subdivided for analysis into a nested hierarchy of four levels (genetic, population

or species, community or ecosystem, and landscape or region) or it can be studied in terms of composition (genetic constituency, species and relative propor-tions in a community, and kinds and distribution of habitats and communities), structure (patterns, se-quence, and organization of constituents), and func-tion (evolufunc-tionary, ecological, hydrological, geologi-cal, and climatic processes responsible for the patterns of biodiversity) Diversity likely enhances sta-bility of the ecosystem, defined as the physiochemical setting associated with a community of living organ-isms in complex, multifaceted interactions Biodiver-sity is one characteristic of an ecosystem, and the sim-plest measure of diversity is the number of types of organisms (usually species or another group of organ-isms in the Linnaean classification system) Alpha di-versity is the number of types of organisms relative to abundance, and beta diversity is a relative measure of how much an ecosystem adds to a region

Species richness measures are typically favored in conservation planning as a proxy for overall level of biodiversity However, there are many definitions of species, and species can be hard to identify no matter what one’s theoretical biases (whether one prefers to explain species change by differing contributions of the evolutionary mechanisms of natural selection, mutation, genetic drift, and gene flow operating slowly and gradually over time or by relatively rapid means during more dramatic environmental shifts) Species exist as ecological mosaics and include a vari-ety of phenotypes that evolve as local environments change The variety of phenotypes within a species is another kind of diversity, named disparity; species

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number and species disparity are not necessarily

cor-related Phenotypes are altered or transformed as a

function of phenotypic plasticity, adaptation, and

mi-gration, but there is no standard means of measuring

and comparing morphological difference within or

between species Which aspects of phenotype are of

interest will again depend on the aims of the

re-searcher

About 2 million species have been described, and

counts of the total number of species range from

5 million to 30 million However, monitored species

indicate that there have been dramatic declines

About 6,200 vertebrate species, 2,700 invertebrate

species, and at least 8,500 species of plants from

around the globe were identified as “threatened” in

2009 in the International Union for Conservation

of Nature (IUCN) Red List of Threatened Species

There is particularly intense interest in identifying

re-gions, called “hot spots,” where a large concentration

of species are experiencing especially high levels of extinctions About 44 percent of vascular plants and

35 percent of vertebrates except fish are found in twenty-five hot spots, representing only 1.4 percent of the Earth’s land surface Most are found in the trop-ics Habitats vary in their distribution of biodiversity, but the environments richest in species are tropical rain forests (primarily because of the impressive num-bers of insects), coral reefs, large tropical lakes, and maybe the deep sea Terrestrial habitats tend to be richest in species at lower elevations and in regions with plenty of rainfall In general, geologically and topographically complicated areas are also likely to have more species

All threatened species are at high risk for becom-ing extinct in their natural settbecom-ings because of human impacts that lead to fragmentation and devastation of habitats as well as the spread of nonnative species, the impact of big-business agriculture and forestry,

Rain forests such as El Yunque Caribbean Recreation Area in Puerto Rico are some of the most biodiverse places on Earth (AP/Wide World

Photos)

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tion, direct use of species, global climate change, and

destructive interference with ecosystem processes

Conserved areas are not enough to stop or reverse the

declines Selection of areas to conserve has been

hap-hazard, and most represent limited ecologies with the

poorest soils, steepest slopes, and highest elevations

Valuing Biodiversity

In the 1950’s, biologists assumed that increasing

bio-logical diversity stabilized ecosystems because any

sin-gle aspect of an ecosystem, if changed, should be less

disruptive the greater the level of complexity In the

1970’s, mathematical modeling of complex systems

confirmed that instability increased with biological

complexity, a view that was favored until the models

proved inadequate to describe all the varying aspects

of living ecosystems Nonequilibrium

(unpredict-able) processes also affected species diversity Thus,

interest continued in the relationship between

mea-sures of biodiversity and productivity, which was the

focus of much experimental research in artificial and

natural settings in the 1990’s However, few simple

as-sociations were found, making the outcome of a

dis-ruption to a particular ecosystem difficult to predict

Some diversity is not evident For example,

biodi-versity is partly determined by genes that may be

somewhat or fully expressed, depending on the

selec-tive demands of local conditions Gene expression is

also sensitive to developmental context as well as

se-lection pressures as the organism survives to

repro-duce The prior history of a lineage (phylogeny) is

also relevant Precipitous population declines can

re-duce genetic variability in a lineage, likely lowering its

flexibility in surviving environmental disturbances

Larger populations are more likely to inhabit more

diverse settings and to accumulate more genetic and

phenotypic diversity Longer-lived (older) systems

seem to accumulate more diversity and are better able

to maintain their integrity

Biological diversity can be assessed in terms of

di-versity among species within an ecosystem, their

vary-ing roles in food chains (trophic) networks, their

biogeochemical cycles, and the accumulation and

production of energy Low species diversity can mean

low productivity when, for example, one compares

deserts and tundra to tropical forests, or high

produc-tivity when evaluating energy subsidized agricultural

systems In addition, greater redundancy of species

with similar roles or functions produces a more stable

system that responds more adaptively to disruptions

The difficulty is that the “roles” and functions of vari-ous organisms within a particular local setting are hard to identify and measure, making the outcomes

of any specific disruptions challenging for planners to predict The stability of a system may mean stability of processes rather than continuity of the same group-ing of species

The Organization for Economic Co-operation and Development advocates the use of marketing strate-gies for increasing the types and levels of biodiversity worldwide There are five economically useful kinds

of biodiversity: direct extractive uses such as foods, plants, and animals of commercial value; direct nonextractive uses, including ecotourism, education, recreation, and extracting and making commercially useful plant products for new medications; indirect uses, as in the case of ecosystems that cleanse air and water, provide flood control, or maintain soil systems; option values or utility for future generations; and ex-istence or bequest values, or how much people are willing to pay to preserve biodiversity Support for bio-diversity will occur if benefits are made explicit and marketable in the global economy

Managing Biodiversity Humans are part of an evolving lineage and are also part of global biodiversity Human population growth and the integration of rural, formerly isolated peo-ples into the global economic system have led to ex-tensive losses of human languages, worldviews, and knowledge about local ecologies and biodiversity No human group should be forced to live on the brink of starvation with high rates of mortality and be ex-cluded from discussions about their region’s biodiver-sity In addition, humans scrambling to survive also have suppressed immune systems and are vulnerable

to epidemic disease

Protection and adequate management of biodiver-sity require that humanity give up the typical short-term, immediate-needs perspective dominated by the most wealthy and politically influential interests and move in the direction of collaboration among diverse interests, including all levels of government, nongov-ernmental organizations, the public, industry, prop-erty owners, developers, and scientists representing academia, government, and industry The planning and associated decision making must include focus

on both public and private lands

Contemporary agricultural systems influence and are influenced by surrounding ecologies less affected

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by human activities Genetically modified plants may

introduce traits that can alter their “wilder” cousins

Agricultural biodiversity has also been declining at

precipitous rates because of reliance on fewer species

as large corporations homogenize and simplify

indus-trial agriculture with reliance on one (monocrop) or

just a few domesticated species All regions are

report-ing declines in mammal, bird, and insect pollinators

This loss of biodiversity in “wild” and “domestic”

ecol-ogies increases the susceptibility of these plants and

animals to virulent diseases that do not stop at

agricul-tural or naagricul-tural boundaries, threatening both

eco-nomic and political stability in affected regions

Conservation

Preservation of species in their natural (in situ)

set-tings involves legislation to protect species, setting

aside protected areas, and devising effective

manage-ment plans, all of which are expectations of the

agree-ment made at the Earth Summit A reserve may

in-clude a less disturbed core surrounded by buffer

zones that differ in the intensity of human use The

designs of reserves are influenced by the research and

theory of the discipline of ecology Larger protected

regions are better than smaller; closely placed blocks

of habitats are better than widely spaced blocks; and

interconnected zones are better than isolated ones

All planning must involve the local peoples living in

or adjacent to the protected regions

Many situations exist in which there is too much

disturbance by humans or the remnant population is

too small to survive under current conditions Thus,

the maintenance of these species in artificial ex situ

(off-site) conditions—such as zoos, aquariums,

botan-ical gardens, and arboretums—under human

super-vision becomes necessary Sometimes captive

colo-nies can be used to introduce species into the wild

Seed banks and sperm preservation are other ways to

conserve genetic diversity, an idea initially pushed by

Nikolai Ivanovich Vavilov in the early twentieth

cen-tury Gary P Nabhan advocates a means of increasing

the biodiversity of local plants and the resulting foods

in a sustainable manner by creating markets

patron-ized by restaurant chefs as well as home cooks for

lo-cally grown, traditional foods Many creative

strate-gies will be required to stop the declines in

biodiversity, which, over time, will most likely increase

the stability and predictability of the Earth’s living

re-sources

Joan C Stevenson

Further Reading

Chivian, Eric, and Andrew Bernstein Sustaining Life: How Human Health Depends on Biodiversity New

York: Oxford University Press, 2008

Cockburn, Andrew An Introduction to Evolutionary Ecology Illustrated by Karina Hansen Boston:

Blackwell Scientific, 1991

Farnham, Timothy J Saving Nature’s Legacy: Origins of the Idea of Biological Diversity New Haven, Conn.:

Yale University Press, 2007

Groves, Craig R Drafting a Conservation Blueprint: A Practitioner’s Guide to Planning for Biodiversity

Wash-ington, D.C.: Island Press, 2003

Jarvis, Devra I., Christine Padoch, and H David

Coo-per, eds Managing Biodiversity in Agricultural Ecosys-tems New York: Columbia University Press, 2007 Jeffries, Michael J Biodiversity and Conservation 2d ed.

New York: Routledge, 2006

Ladle, Richard J., ed Biodiversity and Conservation: Critical Concepts in the Environment 5 vols New York:

Routledge, 2009

Lévêque, Christian, and Jean-Claude Mounolou Bio-diversity New York: John Wiley and Sons, 2003 Louka, Elli Biodiversity and Human Rights: The Interna-tional Rules for the Protection of Biodiversity Ardsley,

N.Y.: Transnational, 2002

Lovejoy, Thomas E., and Lee Jay Hannah, eds Climate Change and Biodiversity New Haven, Conn.: Yale

University Press, 2005

Maclaurin, James, and Kim Sterelny What Is Biodiver-sity? Chicago: University of Chicago Press, 2008 Mann, Charles C Noah’s Choice: The Future of Endan-gered Species New York: Knopf, 1995.

Nabhan, Gary Paul Where Our Food Comes From: Re-tracing Nikolay Vavilov’s Quest to End Famine

Wash-ington, D.C.: Island Press, 2009

Organization for Economic Co-operation and

Devel-opment Harnessing Markets for Biodiversity: Towards Conservation and Sustainable Use Paris: Author,

2003

Primack, Richard B Essentials of Conservation Biology.

4th ed Sunderland, Mass.: Sinauer Associates, 2006

Wilson, Edward O The Diversity of Life Cambridge,

Mass.: Belknap Press of Harvard University Press,

1992 Reprint New York: W W Norton, 1999

Zeigler, David Understanding Biodiversity Westport,

Conn.: Praeger, 2007

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Web Sites

Heritage Canada

The Canadian Biodiversity Web Site

http://canadianbiodiversity.mcgill.ca/english/

index.htm

U.S Geological Survey

Biodiversity

http://www.usgs.gov/science/science.php?term=92

See also: Animals as a medical resource; Biosphere

reserves; Conservation; Environmental degradation,

resource exploitation and; Genetic diversity; Land

management; Land-use planning; Nature

Conser-vancy; Plants as a medical resource; Population

growth; Species loss

Biofuels

Category: Energy resources

Where Found

Biofuels are made mainly from plant material such as

corn, sugarcane, or rapeseed Theoretically, biofuels

can be generated anywhere on Earth where living

or-ganisms can grow

Primary Uses

Biofuels such as ethanol and biodiesel are excellent

transportation fuels that are used as substitutes or

sup-plements for gasoline and diesel fuels Biofuels can

also be burned in electrical generators to produce

electricity Two biofuels are used in vehicles: ethanol

and biodiesel Biogas and methane are used mainly to

generate electricity Biomass was used traditionally to

heat houses

Technical Definition

Biofuels are renewable fuels generated from or by

or-ganisms They can be manufactured from this organic

matter and, unlike fossil fuels, do not require

millen-nia to be produced Since they are renewable, biofuels

are considered by many as potential future substitutes

for fossil fuels, which are nonrenewable and

dwin-dling Moreover, pollution from fossil fuels affects

public health and has been associated with global

cli-mate change, because burning them in engines

re-leases carbon dioxide (CO) into the atmosphere

Using biofuels as an energy source generates fewer pollutants and little or no carbon dioxide In addi-tion, the utilization of biofuels reduces U.S depen-dence on foreign oil

Description, Distribution, and Forms Over millions of years, dead organic matter—both plant and animal organisms—played a crucial role in the formation of fossil fuels such as oil, natural gas, and coal Since the nineteenth century, humans have increasingly depended on fossil fuels to meet energy needs As the supply of fossil fuels has diminished, humankind has begun looking for alternative en-ergy sources Thus, the use of biofuels—including ethanol, biodiesel, methane, biogas, biomass, biohy-drogen, and butanol—is increasing

Ethanol is a colorless liquid with the chemical for-mula C2H5OH Another name for ethanol is ethyl al-cohol, grain alal-cohol, or simply alcohol

Biodiesel is a diesel substitute obtained mainly from vegetable oils, such as soybean oil or restaurant greases It is produced by the transesterification of oils, a simple chemical reaction with alcohol (ethanol

or methanol), catalyzed by acids or bases (such as so-dium hydroxide) Transesterification produces alkyl esters of fatty acids that are biodiesel and glycerol (also known as glycerin)

Methane is a colorless, odorless, nontoxic gas with

Biofuel Energy Balances

The following table lists several crops that have been consid-ered as viable biofuel sources and several types of ethanol, as well as each substance’s energy input/output ratio (that is, the amount of energy released by burning biomass or ethanol, for each equivalent unit of energy expended to create the sub-stance).

Biomass/Biofuel

Energy Output per Unit Input

Oilseed rape (with straw) 9.21

Source: Data from the British Institute of Science in

Society.

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the molecular formula CH4 It is the main chemical

component (70 to 90 percent) of natural gas, which

accounts for about 20 percent of the U.S energy

sup-ply Methane was discovered by the Italian scientist

Alessandro Volta, who collected it from marsh

sedi-ments and showed that it was flammable He called it

“combustible air.”

Biogas is a gas produced by the metabolism of

mi-croorganisms There are different types of biogas

One type contains a mixture of methane (50 to 75

per-cent) and carbon dioxide Another type comprises

primarily nitrogen, hydrogen, and carbon monoxide

(CO) with trace amounts of methane

Biomass is a mass of organisms, mainly plants, that

can be used as an energy source Plants and algae

con-vert the energy of the Sun and carbon dioxide into

en-ergy that is stored in their biomass Biomass, burning

in the form of wood, is the oldest form of energy used

by humans Using biomass as a fuel source does not

re-sult in net CO2emissions, because biomass burning

will release only the amount of CO2it has absorbed

during plant growth (provided its production and

harvesting are sustainable)

Molecular hydrogen (H2) is a colorless, odorless,

and tasteless gas It is an ideal alternative fuel to be

used for transportation because the energy content of

hydrogen is three times greater than in gasoline Also,

it is virtually nonpolluting and a renewable fuel Using

H2as an energy source produces only water; H2can be

made from water again A great number of

microor-ganisms produce H2from inorganic materials, such as

water, or from organic materials, such as sugar, in

re-actions catalyzed by enzymes Hydrogen produced by

microorganisms is called biohydrogen

Butanol (butyl alcohol) is a four-carbon alcohol

with the molecular formula C4H9OH Among other

types of biofuels, butanol has been the most

promis-ing in terms of commercialization It is another

alco-hol fuel but has higher energy content than ethanol

It does not pick up water as ethanol does and is not as

corrosive as ethanol but is more suitable for

distribu-tion through existing pipelines for gasoline However,

compared to ethanol, butanol is considered toxic It

can cause severe eye and skin irritation and

suppres-sion of the nervous system

History

The concept of biofuels is not new People have been

using biomass such as plant material to heat their

houses for thousands of years The idea of using

hy-drogen as fuel was expressed by Jules Verne in his

novel L’Île mystérieuse (1874-1875; The Mysterious Island,

1875) In 1900, Rudolf Diesel, the inventor of the die-sel engine, used peanut oil for his engine during the World Exhibition in Paris, France Henry Ford’s first (1908) car, the Model T, was made to run on pure eth-anol Later, the popularity of biofuels as a fuel source followed the “oil trouble times.” For example, bio-fuels were considered during the 1970’s oil embargo Early in the twenty-first century, concerns about global warming and oil-price increases reignited interest in biofuels In 2005, the U.S Congress passed the En-ergy Policy Act, which included several sections lated to biofuels In particular, this energy bill re-quired more research on biofuels, mixing ethanol with gasoline, and an increase in the production of cellulosic biofuels

Obtaining Biofuels Ethanol is produced mainly by the microbial fermen-tation of starch crops (such as corn, wheat, and bar-ley) or sugarcane In the United States, most of the ethanol is produced by the yeast (fungal) fermenta-tion of sugar from cornstarch Ethanol can be pro-duced from cellulose, the most plentiful biological material on Earth; however, current methods of con-verting cellulosic material into ethanol are inefficient and require intensive research and development ef-forts Ethanol can also be produced by chemical means from petroleum Therefore, ethanol that is produced by microbial fermentation is commonly re-ferred to as “bioethanol.”

In the United States, biodiesel comes mainly from soybean plants; in Europe, the world’s top producer

of biodiesel, it comes from canola oil Other vegeta-tive oils that have been used in biodiesel production are corn, sunflower, cottonseed, jatropha, palm oil, and rapeseed Another possible source for biodiesel production is microscopic algae (microalgae), the mi-croorganisms similar to plants

Methane is produced by microorganisms and is an integral part of their metabolism Biogas is produced during the anaerobic fermentation of organic matter

by a community of microorganisms (bacteria and ar-chaea) For practical use, methane and biogas are generated from wastewater, animal waste, and “gas wells” in landfills Biomass is produced naturally, in the forest, and agriculturally, from agricultural resi-dues and dung

No commercial biohydrogen production process

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