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Tiêu đề Early- and late-onset Alzheimer disease: are they the same entity?
Tác giả P. Tellechea, N. Pujol, P. Esteve-Belloch, B. Echeveste, M. R. García-Eulate, J. Arbizu, M. Riverol
Trường học University of Navarra
Chuyên ngành Neurology
Thể loại Review article
Năm xuất bản 2016
Thành phố Pamplona
Định dạng
Số trang 10
Dung lượng 1,08 MB

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Early and late onset Alzheimer disease Are they the same entity? ARTICLE IN PRESS+Model Neurología 2016;xxx xxx—xxx NEUROLOGÍA www elsevier es/neurologia REVIEW ARTICLE Early and late onset Alzheimer[.]

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www.elsevier.es/neurologia

P Tellecheaa, N Pujola, P Esteve-Bellocha, B Echevestea, M.R García-Eulateb,

J Arbizuc, M Riverola , ∗

aDepartamento de Neurología, Clínica Universidad de Navarra, Pamplona, Navarra, Spain

bDepartamento de Radiología, Clínica Universidad de Navarra, Pamplona, Navarra, Spain

cDepartamento de Medicina Nuclear, Clínica Universidad de Navarra, Pamplona, Navarra, Spain

Received30April2015;accepted14August2015

KEYWORDS

Alzheimerdisease;

Early-onset;

Late-onset;

Neuropsychology;

Neuropathology;

Neuroimaging

Abstract Early-onsetAlzheimerdisease(EOAD),whichpresentsinpatientsyoungerthan65 years, hasfrequently been described ashaving differentfeatures from thoseoflate-onset Alzheimerdisease(LOAD).Thisreviewanalysesthemostrecentstudiescomparingtheclinical presentationandneuropsychological,neuropathological,genetic,andneuroimagingfindingsof bothtypes inordertodeterminewhether EOADandLOADaredifferententities ordistinct formsofthesameentity.WeobservedconsistentdifferencesbetweenclinicalfindingsinEOAD andinLOAD

Fundamentally,theonsetofEOADismorelikelytobemarkedbyatypical symptoms,and cognitiveassessmentspoint topoorerexecutiveandvisuospatialfunctioningandpraxiswith lessmarkedmemoryimpairment.Alzheimer-typefeatureswillbemoredenseandwidespread

inneuropathologystudies,withstructuralandfunctionalneuroimagingshowing greaterand morediffuseatrophyextendingtoneocorticalareas(especiallytheprecuneus).Inconclusion, availableevidencesuggeststhatEOADandLOADare2differentformsofasingleentity.LOAD

islikelytobeinfluencedbyageing-relatedprocesses

©2015SociedadEspa˜noladeNeurolog´ıa.PublishedbyElsevierEspa˜na,S.L.U.Thisisanopen accessarticleundertheCCBY-NC-NDlicense(http://creativecommons.org/licenses/by-nc-nd/ 4.0/)

PALABRAS CLAVE

Enfermedadde

Alzheimer;

Inicioprecoz;

Iniciotardío;

Enfermedad de Alzheimer de inicio precoz y de inicio tardío: ¿son la misma entidad? Resumen LaenfermedaddeAlzheimerdeinicioprecoz(EAIP),definidacomolaquese mani-fiestaantesdelos65a˜nosdeedad,muestraciertascaracterísticasdiferentesdelaenfermedad

deAlzheimerdeiniciotardío(EAIT).Nuestroobjetivofueanalizarlostrabajosmásactualesque

夽 Pleasecitethisarticleas:TellecheaP,PujolN,Esteve-BellochP,EchevesteB,García-EulateMR,ArbizuJ,etal.Enfermedadde

Alzheimer de inicio precoz y de inicio tardío: ¿son la misma entidad? Neurología 2016 http://dx.doi.org/10.1016/j.nrl.2015.08.002

∗Correspondingauthor.

E-mail address:mriverol@unav.es (M Riverol).

2173-5808/© 2015 Sociedad Espa˜ nola de Neurolog´ıa Published by Elsevier Espa˜ na, S.L.U This is an open access article under the CC BY-NC-ND license ( http://creativecommons.org/licenses/by-nc-nd/4.0/ ).

NRLENG-799; No of Pages 10

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Neuropatología;

Neuroimagen

comparanlaclínica,laneuropsicología,lapatología,lagenéticaylaneuroimagendelaEAIPy

laEAIT,paradeterminarsinosenfrentamosadosenfermedadesdistintasoavariantesdeuna mismaentidad.Comoresultado,hallamosconsistenciaenalgunascaracterísticasdiferenciales entrelos2cuadrosclínicos.Fundamentalmente,laEAIPcomienzaconmayorfrecuenciacon unaclínicaatípica;lavaloracióncognitivamuestramayorafectacióndelasfuncionesejecutiva

yvisuoespacialydelaspraxias,ymenorafectacióndelamemoria;laneuropatologíaevidencia mayordensidadyunadistribuciónmásdifusadelapatologíatipoAlzheimer;losestudiosde neuroimagenestructural y funcionalmuestran unaafectación cortical mayor y más difusa, afectandoalneocórtex(especialmenteelprecuneus) Enconclusión,lasevidencias actuales hacenpensarquelaEAIPylaEAITsonvariantesclínicasdeunamismaentidad,queenelcaso

delaEAITseveinfluidaprobablementeporfactoresasociadosalenvejecimiento

© 2015Sociedad Espa˜nola de Neurolog´ıa.Publicado porElsevier Espa˜na, S.L.U Estees un art´ıculoOpenAccessbajolalicenciaCCBY-NC-ND( http://creativecommons.org/licenses/by-nc-nd/4.0/)

Introduction

Alzheimerdisease(AD)isthemostfrequent

neurodegener-ativedisease.Itpresentsclinicallyasprogressivedementia

that predominantly affects episodic memory.1 Age is the

mainriskfactorfordevelopingAD,andinfact,prevalence

ofthediseaseishigherinoldersegmentsofthepopulation

AD is the most frequent cause of dementia and accounts

forapproximately60% ofthetotalcases,whetherbefore2

oraftertheageof65,3theagemarkingthearbitrarylimit

betweenearly-onsetandlate-onsetdementia

In 1907, Alois Alzheimer described the disease now

bearing hisname today in a 51-year-old woman whohad

developed dementia with predominant language

impair-ment andbehavioural changes.4 Foryears, thissyndrome

of early-onset dementia without amnesia was thought to

define AD and distinguish it from senile dementia, which

wascharacterisedbyalateronsetandsymptomsof

amne-sia andattributedto theageing process.However,in the

1960sand1970s,studies showed thatthe neuropathology

underlyingearly-onsetandseniledementiawasthesame;

assuch, both clinical variants are produced by the same

disease,AD.5,6Fromthat timeon,oncetheclinical

crite-riaforanADdiagnosishadbeenestablished,doctorswere

morelikelytorecognisethesenileonsetform7sinceitwas

far more common, while the other ‘atypical’ early-onset

variantwasalmostforgotten.However,anumberofrecent

studieshaveunderlinedtheclinicalheterogeneityofADand

investigateditsunderlyingcauses

This reviewaimstopresent the similaritiesand

differ-ences between early-onset AD (EOAD) and late-onset AD

(LOAD)withregard totheclinical features,

neuropsychol-ogy,neuropathology, genetics, and neuroimaging features

described to date This will serve to clarify if there are

indeed2differententitiesor2clinicalvariantsofthesame

disease

Clinical presentation

The most common initial clinical presentation of AD

is episodic memory loss, which is accompanied by

progressiveimpairmentofothercognitivedomains Never-theless,somepatientspresentalterationsinothercognitive areas and memory remains relatively well preserved AD mayevenpresentasa‘focal’syndromeinwhich the pre-dominantsymptomisapraxiaoranimpairmentoflanguage, visual function, or visuospatialreasoning.This wide array

ofcasesillustratestheheterogeneousclinicalpresentations

of AD, and this results in major challenges and frequent diagnosticerrors.8Somesuch‘atypical’syndromescan eas-ily bemistaken for other entities,suchasfrontotemporal dementia(FTD), whenexecutive or languagedysfunctions aredominant;orcorticobasaldegeneration,whenthereare signsofcorticobasalsyndrome.Assigningthecorrect diag-nosis to ‘focal’ syndromes of AD (especially when ruling out FTD in the differential diagnosis) will involve detec-tinganytemporallobeandposteriorhemispheresymptoms (amnesia,visuospatialdysfunction)ontheonehand,andon theother,verifyingwhetherthefocaldeficitsinthese syn-dromesseemtobelessselective(andprofound)thanthose

in FTD.Here,an exhaustiveneurologicalexamination will revealdeficitsinothercognitivedomainsandthose affect-ingmultiplefunctionalsystemswithinasingledomain(for example,phonology,spelling,andsyntaxwithinlanguage).9

Inlightofthisheterogeneousarray,someauthorshave attempted to facilitate diagnosis by elaborating classifi-cation systems for specific AD subtypes: the typical form (memorylosswithotherdeficits)andthetemporalvariant (with isolated memory loss) are late-onset syndromes In contrast, theleft/language variant (anonfluent aphasia), progressivelogopenicaphasia (with preservedfluency and predominantrepetitiondeficit),right/visuoperceptive vari-ant (including posterior cortical atrophy, which is almost always due to AD) and the frontal/executive variant are early-onsetsyndromes.10 Usingasimilarapproach,another studygroupedADpatientsin3categories:thosewithfrontal lobe dysfunction, very early onset, and a family history; another with primarily posterior deficit (temporoparietal and/or occipitallobes) andearlyonset,andathird group with predominantly temporal lobe dysfunction in elderly patients.11Lastly,currentdiagnosticcriteriaforADdescribe

a typical amnestic presentation and other non-amnestic presentations,includingthosewithpredominantdeficitsin language,visuospatialfunction,andexecutivefunctions.12

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or atypicalclinical presentationsaremore frequently

dis-playedbypatientswithEOAD.8,10,13,14Infact,athirdofthe

patientswithEOADdisplayanatypicalpresentation,

com-paredto6%ofthosewithLOAD.8,11Ontheotherhand,most

studies15—18 haveconcludedthatEOADrunsamore

aggres-sivecourse

Neuropsychology

Numerous articles have compared the neuropsychological

profilesofpatientswithADtodeterminewhethersome

cog-nitivedomainsaremoreaffectedthanothersgivendifferent

agesofonset.Mostsuchstudies19—27indicatesignificant

dif-ferences.Here,weofferasummaryoftheresultsforeach

cognitivedomain

Memory

Ingeneral,studiesshowthatthepatientswiththegreatest

impairmentofthisdomainhave LOAD.16,17,20Furthermore,

memory seems to be relatively well-preserved in the

early stages of EOAD with respect to LOAD.19

Specifi-cally,researchers have observed that LOAD hasa greater

impact on memory, whether of recent events17,20 or of

well-consolidatedinformation,16 althoughsomecasesonly

exhibitimpairedrecognition.21Otherstudies22,23indicatea

qualitativedifferenceinimpairmentwithmemoryloss

pre-dominatinginEOADvsmemoryencodingfailureinLOAD.23

Researchershavealsoobservedpoorertemporalorientation

inthegroupwithLOAD,17,22whichcanbeattributedtothe

accentuatedmemorylossinthisgroupofpatients.22

Language

PatientswithLOADperformmorepoorlyonvisual

confronta-tionnamingtests,suchastheBostonNamingTest.20,21Some

studiesreachedthesameconclusion,butalsofoundthatthe

naming function deteriorated more quicklyin EOAD.24 On

theotherhand,subjectswithEOADscoreloweronwriting

tasks.20Nevertheless,otherstudieshavenotfoundany

dif-ferencesinlanguage,19,25meaningthatthelanguagedomain

is one of the most controversial when comparing these 2

variants

Executive function

DifferentstudiesindicatethatpatientswithEOADwill

per-formmorepoorlyoncomplexattentionandworkingmemory

tasks16,19—21,23andshowmoreerrorsonresponseinhibition

tasks.21

Visuospatial function

Ingeneral,patientswithEOADwillhavepoorerresultsthan

thelate-onsetgrouponvisual-cognitivetasks,19,20,26

refer-ringtoboththeobjectperceptiondomain19andthespatial

perception19,26andconstructiondomains.20,26

Learned motor behaviours

NumerousstudieshaveshownthatpatientswithEOADshow moreseverelyaffectedmotorbehaviours.17,22

Behaviour disturbances

One study compared the prevalence of psychiatric and behavioural symptoms between subjects with EOAD and LOAD andcomparable severity of dementia; it concluded thatthelate-onsetgroupwasmoreaffected Inanycase, thefactthatdysphoriaandapathyarerelativelyfrequent

intheEOADgroupmayberelevant.28

In summary, patients with EOAD perform more poorly

ontasks related towritten language, executive function, attention,visuospatialabilities,andmotor skills,whereas patients with LOAD show poorer performance on tasks involvingepisodicmemoryandvisualconfrontationnaming

Neuropathology

CharacteristicneuropathologicalfindingsinADare extracel-lulardepositsof senileplaquesconsisting ofamyloid beta andintracellularneurofibrillarytanglescomposedof hyper-phosphorylatedtauprotein.Depositionofamyloidplaques begins in the basal regions of the frontal, temporal, and occipitallobesandprogressestoaffecttheprimarysensory areas.Neurofibrillarytangles, ontheotherhand, willfirst affectthe transentorhinalregion andprogresstoward the limbicsystembeforefinallyreachingtheneocortex.29

Thebrain’sdistributionofAD-specificpathologyis corre-latedwiththetypeofclinicalpresentation.Infact,Murray

etal.30 describe3patternsofneurofibrillarytangle distri-butioninAD:thetypicalpatterndescribedabove,another patterninwhichthehippocampusisspared(withmore tan-gles in the cortexthan in the hippocampusand withless hippocampalatrophy),andapredominantlylimbicpattern

In ourstudy, curiously enough, senile plaque density was similarinall3ofthelistedpatterns.Thepatternpreserving thehippocampuswasassociated withearly onset,amore aggressivecourse,andahigherprevalenceofatypical pre-sentations(upto30%).Basedontheseobservations,some researchershavepostulatedthataslightlydifferent patho-logicalcascademaybeatworkinpatientswithanatypical presentation Here, although onset would also be deter-minedbyamyloiddeposits,theformationofneurofibrillary tanglesoccursearlieranditstopographicalpatternisvery different.27

Variousteams have attemptedtoquantifypathological features in AD to compare patients withearly- and late-onsetformsofthedisease.Generallyspeaking,authorshave determinedthattypicalEOADpatientsdisplayagreater den-sityofsenileplaques15andneurofibrillarytangles10,31,32and greaterneuronallosscomparedtotheircounterpartswith LOAD14ortothosewithatypicalEOAD.31Thelatteralsoshow moremarkedimpairmentoftheneocortexthandotypical cases.31ThisseemstoindicatethatEOADhasamore aggres-sive course, but some interpret these findings within the contextof thebrain’s functional reserve; they state that

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Table 1 SummaryofthearticlesconsultedonthesubjectofneuropsychologyinEOADandLOAD.

Study Areaofstudy Results

Gradyetal.65 All NosignificantdifferencesbetweenEOADandLOADfor

anyarea

LOAD:poorerremoterecall

LOAD:poorerrecallofrecentevents,poorerorientation andnaming

Imamuraetal.24 Language EOAD:impairedverbalcomprehensionandnaming Fujimorietal.26 Visuocognitive EOAD:moreimpairmentinattentionand

spatial/constructionalperception Suribhatlaetal.20 All EOAD:poorerwrittenlanguage,attention,and

visuoconstructiveskills;nosubstantialdifferences Kalpouzosetal.23 Working,semantic,and

episodicmemory

EOAD:poorerworkingmemorythaninLOAD;no differencesinepisodicmemory

Snowdenetal.11 All Youngerageofonsetforfrontalneuropsychological

profiles,‘typicalAD’withlossofmemoryand language/arithmetic/spatialskills,andvisual neuropsychologicalprofiles,comparedtoamnesticand semanticprofiles

function;differencesnotsignificantaftercorrectingfor ageandeducationallevel

Toyotaetal.28 Psychiatricandbehavioural

symptoms

EOAD:lowerprevalenceofpsychiatric/behavioural symptoms(delirium,hallucinations,agitation, disinhibition,abnormalmotorbehaviour)comparedto LOADatequivalentdementiaseverity

dysfunction/apraxia,languageimpairment, aphasia-agnosia-apraxiasyndrome,dysexecutive syndrome,andposteriorcorticalatrophy,withless memoryloss

Kaiseretal.21 Language,visuospatial

abilities,executivefunction, andattention,memory

EOAD:poorerfluencywithphoneticcue,more pull-to-stimuluserrorsonvisuoconstructivetasks, poorercomplexattentionandworkingmemory,poorer executivefunction

LOAD:poorernaming,poorerdeferredrecall

apraxias LOAD:poorerorientationandvisualmemory Smitsetal.19 Memory,language,visuospatial

ability,executivefunction,and attention

EOAD:poorerperformanceinvisuospatialability, executivefunction,andattention

LOAD:poorermemory

patientswithLOADwillnotrequiresuchahigh

pathologi-calloadaspatients withEOAD inordertoexhibitclinical

symptoms.32

Theroleofsolubleoligomericformsofamyloidbetain

EOADandLOADhasalsobeenstudied;theseformsmayhave

a more direct effect on the loss of neural function than

doesfibrillaryamyloidbeta.Infact,studiespointtoagood

correlationbetweenthelevelofoligomersand

neurotrans-mitter activity (measured by cholineacetyltransferase).33

They also reveal a distinct pattern of oligomer subtypes

for each group: there is a higher level of pentamers

in the insoluble fraction in EOAD than in LOAD,33 which

indicatespotentialdifferencesinthepathogenesisofeach type

Lastly,whenlinkingneuropathologyfindingswithclinical expression for these 2 patient groups, we cannot over-looktheeffectofconcomitantillness.Inyoungerpatients, the correlation between the amyloid load and the level

of dementia is strong, but this is not the case in older patientsinwhomvasculardiseasemayplayastrongerrole

indementia.34

Insummary,studiesindicatethattheearly-onsetformof

AD is moreextensive, andthat differencesaremore pro-nouncedoutsideofthetemporallobe.35

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Cerebrospinal fluid (CSF) markers

The biomarkers recentlydeveloped for CSFarehelpful in

diagnosingAD,andresearchershaveexaminedthe

possibil-ity that theymay be especiallyuseful for EOAD given its

atypicalpresentation Studiesshow thatlevelsofamyloid

beta,totaltauprotein,andphosphorylatedtauproteinare

abnormalinbothEOADandLOAD,butvaluesaresimilarfor

bothgroups.36,37Ontheotherhand,thereseemstobea

cor-relationbetweenthe(pathologically low)levelofamyloid

betainCSFandthedegreeofbrainatrophyinareasthatare

specifictoeachvariant (especiallytheprecuneusinEOAD

andthehippocampusinLOAD).37AmongEOADcases,there

does seem to bea difference between typical and

atypi-calprofiles;thelatterdisplayahigherleveloftotaltauin

CSF,regardlessofdiseasedurationandthedegreeofclinical

severity.Thisfindingindicatesmoreintensedegenerationin

patientswithanatypicalprofile.38

Genetics

ItcannotbeoverlookedthatADmaybecausedbyan

autoso-maldominantmutationonthePSEN1,PSEN2,orAPPgene.35

Thesecasesaccountforlessthan1%ofallpatientswithAD

andtheyareusuallycharacterisedbyearlyonset35;

never-theless,theymakeuponlyasmallpercentageofallpatients

withEOADandremainoutsideofthescopeofthisarticle

Allele␧4ofapolipoproteinE(APOE*4)isthemost

impor-tant risk factor for sporadic AD Presence of this allele,

whetherheterozygousorhomozygous,hasbeenassociated

with a younger age of onset of the disease.35,39,40

How-ever, various neuroimaging studies coincide in that they

show more pronounced atrophy of the medial temporal

lobe and especially the hippocampus in patients carrying

the ␧4 allele in APOE compared to non-carriers,

regard-less of age of onset of the disease.41,42 There was more

extreme hypometabolism in the medial temporal lobe in

␧4/␧4patientsthaninthosewiththe␧3/␧3genotypewithin

theEOADgroup,butnogenotype-relateddifferencesinthe

LOAD group.43 This is consistent with the greater

preva-lenceofbearersofthisalleleamongpatientswithamnestic

presentation9,35andthelowerpercentageofbearersamong

patients exhibiting preservation of the hippocampus.29 To

summarise, we may therefore state that although the ␧4

alleleislinkedtoyoungerageatADonset,thisonlyoccurs

in patients withtypical ADwho necessarily belongto the

LOADgroup,affectingtheyoungerpatientsinthatgroup.35

On the other hand, some studies have indicated that

APOE*4 determines EOAD progression.17,35 As such,in the

absence of ␧4, progression is quicker for EOAD than for

LOAD;where ␧4is present, progressionis similarinthe 2

groups.35 Likewise, patients with EOAD and the ␧4 allele

havebeenfoundtobeatgreaterriskofdeveloping

myoclo-nias with less tremor; this indicates a different clinical

course,determinedbytheAPOEgenotype,withinthegroup

ofpatientswithEOAD.35

It shouldbeunderstood that,inrecent years,

genome-wide association studies (GWAS) have detected

polymor-phismsincertaingenesassociatedwithLOAD,suchasCLU,

CR1,PICALM,SORL1,BIN1,CTNNA3,GAB2,DNMBP,ABCA7,

TREM2,andTOMM40;thelatterisassociatedwithanearlier onsetofLOADdeterminedbytheAPOEgenotype.44

Structural neuroimaging

Structuralneuroimagingstudies,includingcomputed tomo-graphy (CT) and magnetic resonance (MR) studies, are extremely important for the assessment of patients with cognitiveimpairment,andtheyaremainlyusedtoruleout secondarycauses.Furthermore,MRhasshownhigh sensitiv-ityasameansofevaluatingthecerebralatrophyassociated withneurodegenerativeprocesses;inAD,atrophytypically affectsthemedialtemporalregionandextendsposteriorly

totheparietotemporalandfrontalregions.45

Assessment of overall cerebral volume

Multiple MR studies have calculated the rate of cerebral atrophyin patients withAD andfound it tobegreater in EOAD,withadecreaseof2%to3%oftotalbrainvolumein

ayear46vs0.8%inLOADpatients47(notethatthefirststudy includedbothfamilialandsporadiccases).Therateof atro-phyfor allpatients ingeneralis 1.4%and0.6% inhealthy controls.48 Furthermore,studies indicatethat theatrophy rateincreasesby0.32%peryear2inpatientswithEOAD46;

ontheotherhand,studiesofADmakingnodistinctionsby ageof onsethave notpointed tosignificant accelerations

inatrophyandindicateameanvalueof0.09%peryear2.48

AlloftheabovedescribesEOADashavingamoreaggressive course

Assessment of grey matter

Numerousstudieshave comparedatrophy patternsingrey matter between patients with EOAD and LOAD Frisoni

etal.49foundmorepronouncedoverallatrophyinEOADthan

in the late-onset form (19.5% vs 11.9%) Atrophy primar-ilyaffectedneocorticalareas,especiallytheoccipitallobe

inEOADandthehippocampusinLOAD,andthese topogra-phiescorrespondwiththe2differentclinicalpatterns.Later studieshavereplicatedthesefindingsandattachedspecial relevancetotheprecunealatrophyoccurringinEOAD.50—52

Furthermore,the cortical maps that Frisoni etal.49 com-pared showed that atrophy was very diffuse in EOAD, whereasinLOADitwasdirectedatthetemporallobeand thetemporoparietaljunction.53

We know of only one longitudinal study carried out to date that has compared progression of cortical thinning betweenthese groups; it revealed, for EOAD, more rapid and diffuse atrophy of the association cortices (the left middleand frontalgyri, left inferiorparietal lobe, poste-riorareaoftheleftsuperiortemporalgyrus,leftfusiform gyrus,bilateralposteriorcingulargyri,andprecuneus)and comparativelymore atrophy of the left parahippocampal gyrus in LOAD.54 This pattern of atrophy corresponded to themorepronouncedclinicalimpairmentdisplayedbythe EOADgroupintheareasofexecutivefunctions,attention, andlanguage.54

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One recent study investigatedimpairment in the deep

cerebralnucleiinEOAD andLOAD.55 The differentpatient

groupsdisplaydistincttopographicalatrophypatternsinthe

striate.PatientswithEOADexhibitedmoreextensive

atro-phyofthedorsalstriatum,comparedtotheventralstriatum

in thosewithLOAD The authors’ interpretation wasthat

the changes might have been secondary to loss of

affer-encesofthemedialtemporallobeinthecaseoftheventral

striatum,andof theparietallobefor thedorsal striatum

Another longitudinal study compared the atrophy rate of

subcorticalstructuresbetweenthese2patientgroups.56 It

foundgreaterlossesofvolumeinthecaudate,putamen,and

thalamusin patientswithEOAD thanin thosewithLOAD,

whereaslossesinthehippocampusandamygdalafollowed

similarpatterns.Thesefindingsmayberelatedtothefact

that frontal functions decline more rapidly than memory

doesinEOAD

Assessment of white matter

Studieshavepointedtoapredominantly parahippocampal

patternofatrophyinLOAD,comparedtoamorediffuse

pat-ternofposterioratrophyinEOADthatprimarilyaffectsthe

spleniumofthecorpuscallosumandthedorsal

temporopari-etalregions.Thesewhite-matterfindingscoincidewiththe

topographyof affectedgrey matter, indicating thatwhite

matteratrophyissecondarytoatrophyofthegreymatter.41

Arecentstudyuseddiffusiontensorimaging(DTI)to

com-parewhitematter microstructuraldamagebetween the2

patientgroups.57 Theseauthorsobservedmoresevereand

diffuse atrophy in patients with EOAD, and theydescribe

damagetotheinterhemisphericconnections,limbicsystem

and main associative pathways, and the posterior

cingu-latecortex;LOADwasobservedtoaffectmainlythecorpus

callosum

Insummary,EOADdisplaysamorediffuseatrophypattern

in both grey and white matter, and it affects

neocorti-cal areas (especially the precuneus); atrophy in LOAD is

restricted to the hippocampus Furthermore, progression

seemstobequickerinEOAD,anditextendstosubcortical

areas,mostlytotheputamen

Functional neuroimaging

Functionalneuroimagingtechniquesallowdoctorstoassess

numerousfunctions ofthe brain.Oneofthe most

widely-used techniques is positron emission tomography (PET)

with18F-fluorodeoxyglucose(FDG),whichestimatesneural

activity based on the regional rate of glucose uptake.45

Singlephotonemissioncomputedtomography(SPECT)with

99mTc-HMPAO (99mTc-hexamethylpropyleneamineoxime)

estimatesneuralactivitybyprovidinganimageofregional

blood flow in the brain.45 On the other hand, functional

MRIletsusanalysethebrain’sresponsetospecificstimuli,

which is reflected by the haemodynamic changes in the

brainregionsactivatedbyeachstimulus

PET of glucose metabolism

ThetypicalpatternofADanomaliesinPETstudiesconsists

of hypometabolism in the posterior cingulate, the poste-riortemporoparietalcortex,andtheanteriorregionofthe medialtemporallobes.45 Hypometabolismintheseregions

ismoremarkedinEOADthaninLOADaccordingtostudies employing regionof interest analysistechniques (examin-ingstructuresthatarechosenapriori),andalsoaccording

tovoxel-basedtechniques (analysingthe brainasawhole withnoneedforaprioriselectionofspecificregions).45,58,59

Given the same degree of functional impairment accord-ing to the Clinical Dementia Rating (CDR), the patients withEOADshowedgreaterhypometabolism,butthiscould

be attributed tothe younger subjects’ greater functional reserve Furthermore,the hypometabolismcurve is more pronouncedthanthatoftheCDR,whichindicatesthatthe metabolicdysfunctionprogressesmorerapidly.59

Fig.1showsaPET-FDGstudyfromapatientwithLOAD andanotherfromapatientwithEOAD(lowerrow)

SPECT

ThepatternnormallyseenusingSPECTintypicalADis simi-lartothatdescribedusingPET.45Furthermore,patientswith EOAD exhibitmarked hypoperfusionof theposterior asso-ciative corticalareas, whereas late-onset caseswill show hypoperfusioninthemedialtemporalareas.60

In summary, evidence points once again to more pro-nouncedimpairmentoftheneocortex(parieto-occipitaland even frontal) in patients with EOAD, whereas atrophy in LOADismorelimitedtothemedialtemporallobe

Brain functional magnetic resonance imaging

Functional MRI was used in one study that attempted to characterisefunctional connectivitypatternsin EOAD and LOAD.61TheEOADgroupdisplayeddecreasedconnectivityof thedorsolateralprefrontalnetwork(DLPFN,relatedto exec-utive function) andincreased connectivity in theanterior temporal network (ATN, related to declarative memory); patients with LOAD exhibited the opposite pattern The authorslinkedtheirfindingstothedistincttopographiesof neural damage (decreased connectivity) in each of the 2 groups, andthefactthatdamagewouldgiveriseto com-pensatorymechanisms(increasedconnectivity)

PET

Therecentdevelopmentofradiotracersabletobindto amy-loidprotein hasmade it possible toevaluate AD features

in vivo.45 Most of the studies performed using Pittsburgh compound B (PiB) in patients with EOAD and LOAD did notrevealsignificantintergroupdifferences.55,62 According

to theauthors, the discrepancy between PET-FDGimages (which differ significantlybetween groups, asmentioned) and PET-PiB images (which are similar in both groups) indicates that other pathological processes, such as the formationofneurofibrillarytangles,neuroinflammation,or

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Figure 1 Positron emission tomography (PET) studies with florbetapir (upper row; images fused with CT) and 18 F-fluorodeoxyglucose(FDG; lowerrow)inpatients diagnosedwithAlzheimerdisease (A)Womanaged 79diagnosedwithtypical Alzheimerdisease.ThePETstudywithflorbetapirgivesapositiveresultforcorticalamyloidplaques.ThePET-FDGstudyindicates hypometabolismintheposteriorassociationcortex,predominantlyontheleftside.(B)Managed57diagnosedwithleft/language variantAlzheimerdisease.ThePETstudywithflorbetapirgivesapositiveresultforcorticalamyloidplaques.ThePET-FDGstudy indicateshypometabolismintheposteriorassociationcortex,predominantlyontheleftside

amyloid oligomers (mentioned previously and not visible

withPET-PiB),maybemoreinvolvedthanfibrillaramyloid

deposition in eliciting metabolic changes.62 Nevertheless,

Ossenkoppeleetal.63 observed agreaterconcentrationof

theradiotracerintheparietalcortexofpatientswithEOAD

andarecentstudyshowedthatpatientswithEOADdisplayed

more amyloid deposition in the basal ganglia, thalamus,

leftsuperiortemporalcortex,andleftcuneus.64 Thisisto

beexpectedbasedonthePET-FDGmetabolicneuroimaging

studiesthatwehavealreadydescribed,anditwouldfitthe

ideaofapathologicalcascadeinwhichdepositionof

amy-loidbetawouldprecedemetabolicdysfunction.Itremains

tobe seen what would causeamyloid depositsin certain areasandnotinothers.63

Fig.1alsoshowsaPETstudyofamyloiddepositsinone patientwithLOADandanotherwithEOAD(upperrow)

Conclusion

EOAD and LOAD have many traits in common, but they alsoexhibitnumerous differences(Table2).The factthat they display the same pathological process, which is the only defining diagnostic criterion for AD, obliges us to

Table 2 GeneralcharacteristicsofEOADandLOAD

Age at onset 65yearsandolder Youngerthan65years

Form of onset Amnestic Non-amnestic(visuospatialdysfunction,apraxias)

Progression Slower Faster

Neuropsychology Poorermemory Poorerexecutivefunction,visuospatialskill,and

motorskill

Pathology findings Senileplaquesand

neurofibrillarytangles

Senileplaquesandneurofibrillarytangles,with betterpreservationofthehippocampus

Biomarkers in CSF LowerlevelofA␤42and

increaseintauandP-tau

LowerlevelofA␤42andincreaseintauandP-tau

APOE genotype Favouredby1or2␧4alleles Favouredbyabsenceof␧4alleles

Neuroimaging

StructuralMRI Hippocampalatrophy Frontal/temporoparietalatrophy

PET-FDG Decreasedmetabolismin

medialtemporallobe

Decreasedmetabolismintemporoparietalcortex PET11C-PiB Increaseduptake Increaseduptake(moreinparietalzone?)

Modified from van del Flier et al 35

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conclude that they are technically variants of the same

disease.Nevertheless,thesubstantialdifferencesbetween

these variants, which we have listed in this study, raise

the question of whether anatomical pathology findings

alone should constitute what defines a nosologicalentity

when it comes to classifying these neurodegenerative

diseases

In addition, neuropathology and neuroimaging findings

pose interesting questions regarding whether or not the

underlying pathophysiologicalprocess is thesame in both

forms.Perhapsthedifferencesbetweentheseclinical

vari-antscanonlybeexplainedbythefactorsinvolvedinageing

Onthe other hand, theseeminglymore aggressivecourse

ofEOAD calls for furtherinvestigation ofwhy thedisease

wouldexhibitdifferentbehaviourinsomecasesandnotin

others

Insummary, EOADmorefrequentlyexhibits anatypical

early clinical course; furthermore, patients will be more

affected in the areas of executive function, visuospatial

function, and motor skills, and display less memory loss

EOADcasesshowagreaterdensityofamyloidplaquesand

amorediffusedistributionpatternthanthatseeninLOAD

Lastly,EOADcaseswillshowagreaterextensionofatrophy

witha more diffuse pattern and more rapid progression;

these tendencies arealso observed using PET and SPECT

functionalimaging.Ontheotherhand,theleastconsistent

resultsareprovided byPETstudies evaluatingthe brain’s

amyloidload

Conflicts of interest

Theauthorshavenoconflictsofinteresttodeclare

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