However, there are so many fungi are waiting for discovering in Vietnam in genretal, and in Xylaryaceae family in particular, the largest family known in Ascomycota.. In [r]
Trang 1T4p chi Khoa hsc DHQGHN Khoa hsc Tu nhi6n vA C6ng nghQ, Tip 29, Sd 2 (2013) 38-43
B6 sun gba lodi n6m trii h9 Xylariaceae
cho khu hO ndm Viet Nam
D6 Dric Qu6, Ducmg Minh Lam*
Trudng Dqi hoc Su pham Hd NAi, I36 Xudn Thily Str., Cdu Giiiy Dist., Hd N|i, Vi€t Nam
Nhfn ngdy 14 th6ng 3 ndm 201 I Chinh sta ngiry 14 th6ng 8 ndm2012; ch6p nhfln tldng ngiy 07 thang5 nlm 2013
Ttim tit- K6t qui nghidn criu t4i rirng nguy6n sinh Mudng Ph[ng, tinh DiQn Bi€n ndm 2010 chring t6i de b6 sung th6m 3 lodi mdi cho khu hO ndm d ViCt Nam Trong nghidn cuu niy, 3 lodi mdi
thuQc chi Annulohypoxylon (A nitens, A purpureonitens- vd A; stygium) dugc ghi nhdn vd mi6u td, dua s6 lodi thuQc chi ndy d Viet Nam l6n 8 lodi trong t6ng s6 41 lodi hiQn dugc ghi nh{n tr6n thil
gidi Chring t6i da ti6n hdnh x2y dung kh6a ph6n lo4i diin lodi dlra vdo c6c df,c tli6m hinh th6i cia
8 lodi tr6n.
Tii kh6a: Annulohypoxylon,Xylariaceae, Mudng Phdng, DiQn Bi6n, NAm tui.
1 MO tIAu
N6m trii h9 Xylariaceae c6 khoing 3000
, A,
lodi, ld ho c6 sd lugng lodi l6n nhdt trong bQ
n6m trii Xylariales, ngdnh ndm tii Ascomycota
Di5n nay tr€n th6 gdi de x6c tlinh dugc 75 chi
thuQc hg ndy Chring dugc phdn bd o hAu h6t
c6c hQ sinh th6i tlugc nghi€n c(ru trdn tr6i d6t,
nhrmg tf,p trung voi d0 <la d4ng cao d nhirng
vung nhiQt ddi [].
PhAn lcrn n6m trii h9 Xylariaceae cu trf tr0n
g5 mgc dir c6 mQt s5 c6c lodi vi du nhu
Hypoxylon terricola vd Xylaria carpophila t\m
., t
th6y trdn rric Ngugc lai chi Poronia vd chi
Podosordaria Wawelia lgi sting tr€n phAn, mQt
A ,tt
-sd c6 khd ndng gdy bQnh bQnh th6i 16, th6i cdy
vd mtic suong 16 Xylariaceae ld ddi tuqng r6t
' T6cgiit li€n hQ DT: 84-984150666.
E-mai I : quedoduc@gmail.com
ttit cho viQc nghiOn ctu enzim phdn giAi 96 do tlac tti6m sinh trucrng trdn co ch6t g5, m6t s6 lodi n6m trii thuQc chi Xylaria c6 kJrd ning ph6n gi6i 70-80 % lignocellulose trong 16rpng12,3) Ndm 2005 Yu-Minh Ju vd Jack Rogers t6ch
ra tri chi Hypoxylon mQt s6 lodi c6 sg kh6c biQt
vd mat sinh hgc phdn tri vd c6 m6t sO aac Aiem
Annulohypoxylon, ban dAu chi ndy mdi c6 33
lodi [5] DiSn nay c6c nhd khoa hoc cld c6ng b6 clugc 4l lodi thudc chi Annulohypoxylon tr€n
th6 gioi D6 ld kt5t qui tlSng ghi nhdn, tuy nhiOn
ngudi ta u6c chimg vin cdn khodng 50 - 60 lodi chua tlugc tim th6y o chi ndy
O Vigt Nam chi m6i c6ng b6 vd md td dugc
1l chi vdi 5l lodi thu6c h9 ndy [4] Trong il6 c6
cohaerens; A annulatum; A multifurme; A. moriforme vd A pyrifome) Trong qu6 trinh thu 38
Trang 2D.D QuAi D.M Lam lTap chi Khoa hgc DHQGHN, Khoahpc Trt nhiAn ztd C6ng nghQ, TQp 29, Sd2 (207il 3843 39
thap vd nghidn cuu c5c mdu n6m t4i rung
nguy€n sinh Mulng Phdng - DiQn Bi€n, chirng
t6i de ph6t hiQn thdm 3 loii n6m trii thuQc chi
ndy d6 ln (Annulohypoxylon nitens; A
purpureonitens vd A stygium), dua sti loei
trong chi nAy c6 mflt tai ViQt nam l6n 8 lodi
Trong c6ng b6 ndy, 3 lodi m6i dugc bd sung
vdo danh s6ch c6c lodi n6m tfi o ViQt Nam
clugc mi6u td chi tii5t Kh6a phAn lo4i t6i lodi
trong chi Annulohypoxylon c6 mit d ViQt Nam
clugc x6y dqng
2 Nguy6n tiQu vir phuong ph6p
2.1 Nguy€n liQu
C6c m6u n6m trii thu thap ctugc t4i rimg
nguy€n sinh Mudng Phdng, DiQn Bi€n ndm
2010; Chc m6u nAm trii thuQc cht Annulohypoxylon
ttugc thu th{p tir rimg nguy€n sinh Mudng
Phdng - DiQn Bi€n
C6c h6a ch5t sri dung trong nghiOn cr?u ndy
ct6u tlat mftc ti6u chuAn ph6n tich (Merck,
Sigma) C6c thit5t bi nghi€n criu dim bdo <10
chinh x6c cao (kinh hi6n vi soi ni5i Zeiss; kinh
hii5n vi quang hgc Olympus CX41 c6 gin m6y
6nh k! thuat s5; panh si6u nho Thgy s!,
Melzer's cta Merck-Dtc ) .
2.2 Phuong phdp
Phtmng phap thu thQp vd bdo quan mdu pl.
.MAu n6m trii tlugc thu thdp niQt c6ch ngSu
nhi0n trong c6c l*ru.*g Arn t4i Muong Phdng,
DiQn Bi6n vd tlugc d6 vdo c6c tui grdy v6 trung,
ctdm bio git Am cho m6u trong thdi gian v{n
chuyi5n vC phdng thi nghiQm (2 ngdy) Tai
phdng thi nghiQm, c6c m6u dugc chua phdn tich
ngay tlugc de kh6 tg nhiOn Thdi gian phAn tich
mlu t4p trung trong khodng I tuAn, sau tl6 c6c
miu dugc s6y khd vd bdo quin ldu ddi
Phrong phdp phdn tich mdu [2]
Chung tdi 5p dgng phuong ph6p nghi€n cuu
hinh th6i so s6nh fiong qu6 trinh nghi€n cuu vi
phAn tich mdu Anh trii bio tt vd bdo tri clugc chpp vdi vat kinh dAu (x 100), ch6t nAn vd th6
qui tluo c chup vdi kinh hip soi ntii DAy cflng ld phuong ph6p truydn thiing clugc sri dgng tr6n
th6 gioi vd ViQt Nam tir trudc t6i nay trong nghi€n cr?u phdn lo4i n6m
f X6t qufl vh thio lu$n
3.1 Ddc diiim mi€u td chi tt€t hi€n vi ndm fii
Annulohypoxylon o ViQt Nam
Annulohyporylon nitens (Ces.) Y.M Ju &
J.D Rogers(2005)
Basionym Rosellinia nitens Ces., Atti Accad Sci Fis 5:13 1872 Hypoxylon nitens (Ces.) Y.-M Ju & J.D Rogers, A Revision of
the Genus Hypoxylon, p 220 1996
Miu n6m trong nghiOn criu thuQc lodi ndy ld m5u n6m MP0l03
Dia tli6m thu mdu: O tga dO N
21027'02.523, E 1030 09'54.421, tr6n rlQ cao
1024 m so vdi mlrc nu6c bi6n, trong nmg'
nguy6n sinh Mudng Phdng, xd Mudng Phdng, huyQn DiQn Bi6n, tinh DiQn Bi€n
Thdi gian thu m6u: 3l7l2ol0
M6i truong s6ng: tr€n thdn g5 dang trong
giai tlo4n dAu cria qu5 trinh phAn hriy, tr€n bA
m{t v6 cta cdy, trong <lidu kiQn <tri 6nh s6ng, dQ
Am kh6 cao.
Chtit nin phing dgt, hinh gOi toa mgc thdnh
c6c cpm to nhd kh6c nhau, kich thudc 0.2-l.l
cm chi6u ddi x 0.2 - 0.6 cm chi6u ngang, b6 mflt mdu den sAn siri, nhin 16 c5c thi5 quA, chim
khoing l/4 trong g5, phdn fmg v6i dung dich
KOH 10% cho mAu xanh 16 cAv dtin mdu xanh
dam.
Trang 340 D.D QuC D.M Lam lTqp chi Knahoc DHQGHN, Khoahoc Ty nhi€n od C6ng nghQ, Tqp 29, Sd2 Q013) 3843
Thii qud hinh cAu, mAu den c6 tludmg kinh
330 - 550 pm, v6 cimg, gidn d4ng cacbon.
Hinh l Ddc tli6m hinh th6i m6u n6m MP0l03
(Annulohyporylon nilens).
a Chiit nin; b Bd mdt chh nin; c Ldt cit ngang chdt
nin; h Tii Bdo tu d; e Phan ung miu chdt nin voi
KOH I0%; f.V6 bdo t* bi tdch tong KOH l0o%; c.
Rdnh ndm; h Dinh ttii bdo t*; Ti lQ: a 1000 pm; b,c,
e: 500 p4 d: l0 pm f,g, h : 7 pm
.:
L6 miQng c6 nhri cao hon bO mpt ch6t n6n,
dugc bao quanh boi mQt mflt phing hinh clia,
tluong kinh 240 - 260 pm
Tui bdo /ri hinh tru dei, chria 8 bdo tri, v6
.!
clcrn trong su6t voi tdng chi€u dei 100 - 120 pm
x 4 - 5 pm chi€u ngang, phAn c6u truc mang bdo
tri ddi 60 - 65 pm, phAn cut5ng kh6ng mang beo
tt ddi 40 - 45 pm Dinh tiri bdo tri bit mAu xanh
v6i thu6c nhuQm Melzer's, hinh tlia" kich thudc
0.5 - 0.7 pm chidu cao x | - 1.2 pm chidu
ngang.
Bdo tti tili hlnh elip v6i mAu ndu tl6n ndu
nh4t, kich thu6c 6.5 - 8 pm chi6u ddi x 3 - 3.5
li ) I r I
pm chi6u ngang, rdnh mdm ch4y dgc gdn h€t
chi6u ddi bdo tu, vd bdo tu bi t6ch trong dung
dlch KOH lO% ((t gAn dAu cria v6 c6 mQt di6m
dny l€n)
Dga vdo c6c chi ti€u phdn lo4i di5n lodi vd
c6c dflc di6m, hinh th6i, c6u truc, kfch thu6c tui
bdo tu vd bdo tri cta mdu n6m MP0l03, ta theo
lfi6a phdn lo4i <ta5n lodi trong chi Annulohypoxylon dd cdng bd cria Yu-Minh Ju
vd Jack Rogers Chring t6i de x6c dinh clu-o c
m6u n6m ndy thuQc lodi Annulohypoxylon nitens.
Dac tti€m itflc tnmg nhAt cria lodi ndy ld
ch6t nAn v6i nhirng thi5 qui x6p thdnh chtrm, hinh g5i nhd, c6 mAu den vd kfch thu6c bdo tri
dditir6-8pm.
Lodi ndy tl6 tlugc tim thAy d Indonesia,
Malaysia, M!, Mexico vd d ViQt Nam trong nghi€n cftu ndy Ddy lA lodi mdi dugc c6ng b6
Annulohyporylon purpureonitens ; Y .M Ju
& J.D Rogers (2005)
Basionym Hypoxylon purpureonitens
Y.-M Ju & J.D Rogers, A Revision of the Genus Hypoxylon, p 224 1996
M6u nAm trong nghiOn cr?u thuQc lodi ndy
li miu n6m MP0095
Dia <ti6m thu m6u: d tea dO N 21026'59.135, E 103009'49.320, tron dQ cao
1010 m so v6i mgc nudc bi6n, tgi rung nguydn
sinh Mudng Phdng, xd Muong Phdng, huyQn DiQn Bi6n, tinh DiQn Bi6n
Thoi gian thu m6u: 3 /7 l2OlO
Mdi truong sSng: O tr6n thAn g5 dang
rl trong giai <lo4n tl6u cria qu6 trinh phAn hty,
nim tr€n bO mit vo cAy, trong did:u kiqn kh6 t6i,
dQ Am cao.
Chiit nin phing dst hinh g6i kicn thu6c 9.0
- 10 cm chi6u dii x2.5 - 4.0 cm chidu ngang,
bA mat c6 mAu ndu x6m d6n ndu den nhin 16 c6c th6 qud, chim 1/4 trong g5, phdn rrng mAu vdi
dung dich KOH l0% cho mAu nAu xiim nh4t.
ThA qud hinh cAu, mAu den c6 kich thudc
615 - 660 pm, v6 cimg gidn d4ng cacbon.
Trang 4D.D Qu4 D.M Lam /Tap chi lQtoahqc DHQGHN, Khoahgc Tt nhiAn oi C6ng nghQ, Tfip 29, Sd2 (2073) 38-43 41
Hinh 2 D{c tli6m hinh thei m6u n6m MP0095
(A nnulo hyp o xy I o n p urp ur e o n it en s ).
a Chfu nin; b BO mdt chfu nin; c Ldt cfu ngang chdt
nir; d Trii bdo tv: e.Phan ung miu chdt nin voi KOH
10%; f Dinh tii bdo t*: g Rdnh mdm; h V6 bdo tu bi
tach trong KOH l0%; Ti lQ: a c, e: 1000 pm; b: 500
pm; d: l0 pm f: 4 pm; ,g, h: 7 pm.
-:
L6 miQng c6 nhri cao hctn b€ m|t ch6t ndn,
clugc bao quanh boi mOt ving phing hinh dia,
clucrng kinh220 - 265 pm
Tili bdo lri hinh trg ddi, chr?a 8 bdo tti, vo
,t
don trong su6t voi tdng chi6u ddi 120 - 130 pm
x 4 - 5 pm chi6u ngang, phAn c6u trfic mang bdo
tft ddi 55 - 65 pm, ph6n cu6ng kh6ng mang bio
tfr ddi 60 - 65 pm Dinh trii bdo tu b6t mAu xanh
v6i thu6c nhuQm Melzer's, hinh ctTa vdi kich
thuoc 0.5-0.8 pm chi€u cao x l -1 5 pm chi€u ngang.
Bdo ttr trii hinh elip kh6 tliu, mAu vdng ndu
vdi kich thudc 6 -7 pmchidu ddi x 3 - 3.5 pm
chi€u ngang, c6 rdnh mdm ch4y dqc g6n h€t
chi€u ddi bdo tft, vo bdo tu b! t6ch trong dung
dich KOH l0% (ir gAn dAu cria v6 c6 mQt di6m
ddy l€n)
Qua nhfrng ilac di€m mi€u tA hi€n vi vd
hinh th6i, chring tdi de ti6n hdnh tra theo kh6a
phAn lo4i <t.5n lodi <16 cdng b6 cia Yu-Minh Ju
vir Jack Rogers (1996) [6]vd de x6c tlinh clu-o.c
miu n6m MP0095 ld lodi Annulohypoxylon
purpureonitens Loiti ndy c6 r6t nhi6u di€m gAn
vdi loAi Annulohypoxylon nitens v6 ci kich
thudc trii bdo tu, bio tu vd l5 migng Tuy v4y
sg kh6c uir5t ve miu sic ch6t ndn khi truong thdnh, phin img mAu cria ch6t ndn vdi dung
dich KOH l0%o, ciung nhu kich thudc cria thi5
qud li c6c dflc di6m dO c6 thd phdn biQt vd hinh
th6i gitia 2 lodi n6m ndy Trudc d6y c5c nhd khoa hgc dd x6p 2lodi ndy vdo ldm mQt d6 le
Hypoxylon nitens, nhmg nIm 2005 Ju vd cQng
su dd tach ra thdnh 2 lodi ri€ng nhd c6 nhirng
<tac tti6m kh6c bi-Ot vC hinh th5i cfing nhu trinh
ts DNA [5].
Lodi niy dd dugc tim th6y d Brazil, Mexico
vd Vi€t Nam trong nghi€n criu niy Ddy ld lodi m6i cdng b5 O viet Nam
Annulohyporylon stygium (Ldv.) Sacc;
Y.M Ju & J D Rogers (2005) Basionym Sphaeria stygia Le'v., Ann Sci
Nat Bot, se'r III, 5:258 1846 Hypoxylon
stygium (Le'v.) Sacc., Syll Fwrg.I, p379.1882
MAu nAm trong nghiOn criu thuQc loAi ndy ld m6u n6m MP0079;
Dia di€m thu mdu: d tea d0 N 2102j'02.123, E 1030 og'50.122, tr€n rlQ cao
1011 m so v6i myc nu6c bir5n, t4i rimg nguy6n
sinh Mudng Phing, xd Muhng Phing, huyQn DiQn Bi0n, tinh DiQn Bi6n
Thoi gian thu m6u: thdng 10/ 5 l2OlO M6i tnrcrng s6ng: M6u n6m thu dugc trOn
thdn g5 dang trong giai cto4n clAu ph6n hty,
trong cli6u kiQn dQ Am cao, rim rqp vdi nhi€u
cdy br,ri mgc bao trirm
Chiir n€n hinh g6i voi kich thudc 0.7 - 5.5
cm chi6u ddi x 0.4 - 1.5 cm chi€u ngang, bd mflt miu den xen l6n nhirng d5m tring nhd li ti
v6i c6c th6 qud nhin k*r6 16, chim ll4 trong 96, phdn ung mdu v6i dung dich KOHI0% cho mAu xanh h c|'y
Thii qud hinh cdu d6n hinh tru cAu kich thu6c clu&ng kinh 330 - 440 pm;250 -300 pm
chi6u ngang x 300 - 450 pm chidu cao, c6 mAu den, vd d4ng cacbon cimg, gidn
Ld miQng hinh dia duong kinh 175 - 2OO
pm, c6 nhri cao hon b€ mflt ch6t nAn
Trang 542 D.D Qu{, D.M Lam lTap chi Khoahoc DHQGHN, Khoahgc Tt nhiAnod COng nghQ, TQp 29, Sd2 (20Lil 3843
Hinh 3 Ddc <litim hinh th6i mau nAm MP0079
(Annul o hy p ory I on s ty gium).
a Chdt nin; b Bi m4t chdt nir; c Ldt ch ngang chdt
nin; d Tili bdo tn e phan ing miu chdt ninvai KOH
l0%; f Rdnh mdn: g V6 bdo tu bi tdch trong KOH
l0%: h Dinh tili bdo tu: Ti l€: a c, e: 1000 pm; b: 500
pm; f: 4 pm; g: 10 pm d,h : 7 pm
Tili bdo rri hinh hu ddi, chr?a 8 bdo tri, vo
tlon trong su6t v6i t6ng chidu dei 70 - 80 pm x
, i.x
3.5 - 4 pm chi6u ngang, phdn cdu tr0c mang bdo
ffi dni 50 -55 pm, phAn cu6ng kh6ng mang bdo
tir dei 20 - 25 pm, c6u trtic clinh bit mAu xanh
nh4t v6i thu6c nhuQm Melzer's , hinh fra v6i
kich thu6c 0.5 - 0.8 pm chiAu cao x I - 1.2 pm
r l
cnreu ngang.
Bdo t* ttii hlnh elip diu, don bdo, mAu vdng
n6u vdi kich thu6c 6 - 7 pm chi€u ddi x2.5 - 3
pm chiAu ngang, c6 rdnh mAm ddi gAn hi5t bao
tri, v6 bdo tu bi tach trong dung dich KOH l0%
(d gAn ctAu cta v6 c6 mQt <ti6m ddy l6n)
Ddy ld lodi c6 nhfing tlflc tli€m ri6ng biQt r6t
d6 nhan th6y nhu ch6t ndn hinh g5i, phing det,
kich thudc bio tri ngin khoing6 - 7 pm, dudng
kinh l5 miQng nhd hon 200 pm Li lodi it gap
^ .L ,
trong c6c cdng b6 truoc cl6y.
Chring t6i de ti6n hdnh tra kh6a phdn logi
cla5n lodi vd dd x6e tlinh tlugc miu n6m ndy lA
loii Annulohyporylon stygium Lodi ndy c6
a I
quan hQ rilt gdn vdi Annulohyporylon
atroroseum, tuy nhi6n sU kh6c biet h ch6t n6n
khi trudng thdnh d Annulohypoxylon atroroseum c6 mAu h6ng, vdng quanh l5 miQng
c6 mAu den cdn & Annulolrypoxylon stygiumkhi
,l trudng thdnh c6 mAu den b6ng
Lodi ndy dd dugc tim thlry Indonesia,
Malaysia, Mexico, Dii loan, Th6i Lan vd o ViQt
Nam trong nghidn cftu ndy DAy ld lodi m6i lAn
tlAu tlugc cdng bd o Vi€t Nam
3.2 Kh6a clinh loqi tdi lodi cita chi Annuloltyporylon dd dtqc tim thay d Vi€t Nam
Sau khi nghi€n cftu c6c ttac dii5m hinh thrii cria c6c lodi thuQc chi Annlohypoxylon d ViCt
Nam, chfng tdi de xAy dpg kh6a cfinh lo4i cilc loii thuQc chi ndy vd m6 td chc lodi ituqc b6
A
sung cho khu hQ n6m d Vi6t Nam
1 L5 miQng c6 nhri, kh6ng c6 virng bao quanh hinh tlia 2
7
1 L6 miQng c6 nh[, du-o c bao quanh bdi mQt lop phing hinh dTa 3
2 Bdo tri n6u (9 - 11 x 4 - 5 pm), rdnh mAm
, .]
thdng chay dei hCt chi6u ddi bdo
tri 4 cohaerens
2 Bdo tri mAu nAu nh4t dt5n mAu nAu 8.5 - 12 x
3.5 - 5 pm; vdi rdnh mAm ng6n hon chi6u dii
bdo tu A maltiforme
3 ETa xung quanh 15 miQng c6 duong kinh
0.1-0.2 mm; Bdo tft mdu n6u, 5 - 7 x2 - 3 pm1'
Rdnh mim nhin r6t 16, nim tr6n c4nh phing
ri
3.DTa xung quan 16 miQng c6 dudng kfnh 0.2
-0.5 mm; Rdnh mAm nhin r5, nim tr6n c4nh l6i cria bdo tu 4
4 Chdt ndn khi hucmg thdnh c6 miu n6u clfm
titin clen nh4t 5
4 Chitt nAn khi trudng thinh c6 mAu ndu x5m tl
tl6n tlen b6ng, 6
5 Chit ndn mgc thdnh cgm, hinh gi5i, hinh b6n
cffu, hoic hinh gdi t6a, v6ith€ qui mgc thdnh
Trang 6D.D Qu{, D.M Lam lTqp chi Kroahoc DHQGHN, Khoahgc Ty nhi€nad C6ng nghQ, Tqp 29, Sd2 (2013) 3843 43
u nh6 dE nh{n th5y, mdu tten; bdo t& 6-9 x
254 pm; 4.morfurme
5.Ch6t n6n hinh b6n cAu, khdng nhin rd c6c p
mAu nAu den, mAu lgc vdng; s6ng tr€n cdy
:'
s6i, cAy d6 d virng nhiQt tl6i bdo tri 7.5-10.5
(-11) x 3.5-5 (-6) pm .A onnulatum
6 Ch6t ndn phdn fng mAu vdi dung dich KOH
10% cho mAu hoi lUc; bdo tt 6.5-10 x 34.5
; , .^
pm; xudt hiQn o nhi6u noi vd o virng nhiQt
ddi A, nitens
6 ChAt ndn phAn r?ng mAu v6i dung dich KOH
i0 % cho mAu mAu d6 tia, mAu ndu nhat; bdo
tft 6.5 - l0 x 3 - 4.5 pm A purpureonitens
4 K6t tugn
Trong thdi gian nghi€n cr?u ngin, sti lAn thu
;,
thflp mdu m6i chi lir 2 l6'n nhtmg chring tdi de
ph6t hien <lugc 3 lodi m6i ghi nhfln lAn dAu ti€n
o Viet Nam trong chi Annulohyporylon K6t
qud nghiOn criu ndy tld ttua t6ng s6 lodi tong
chi ndy o Viet Nam l€n 8, trong s5 +t loai da
^ ,A ^ 4
c6ng bd tr6n th6 gicri K6t qud nghi€n cr?u cdn
cho th6y mirc ctQ cta d4ng cao cria n6m tui trong
rimg nguyCn sinh Mudng Phdng, DiQn Bi6n
Loi cim mr
C6c ktit quA cta nghi€n cftu ndy duo c thgc
hiQn v6i sg tdi trg kinh phi tir de tdi kfroa hgc
c6p BQ Gi6o dpc vd ddo t4o md sO
gZOOg-tZ-245 do TS Duong Minh Lam ldm chri nhiQm.
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Three more ascomycetous species of family Xylariaceae
newly recorded in Vietnam
Od Oric Quti, Duong Minh Lam
Hanoi National (Jniversity of Education; 136 Xudn Thny Str., Cdu Giiiy Dist., Hanoi, Vietnam
Abstract Studies in fungal diversity in Vietnam have long history but there have been few publications on ascomycetous groups Some publications are leading research on this group of fungi taken occured by Trinh Tam Kiet and Ngo Anh However, there are so many fungi are waiting for discovering in Vietnam in genretal, and in Xylaryaceae family in particular, the largest family known
in Ascomycota In our research on fungal diversity in Muong Phang, Dien Bien, in 2009, we already
published 3 newxylariaceous species Inthis article,3 species of Annulohyporylon genus (1 nitens,
A purpureonitens vd A stygium) are newly recorded in Viebram and described in details
Dichotomous keys to existing species of Annulohypoxylon in Vietnam are developed
Keywords: Annulohypoxylon,MuongPhang, Dien Dian, Xylariaceae, Ascomycetes