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Targeting TKI resistance in non - small cell lung cancer patients caused by secondary EGFR T790m mutation

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Non - small cell lung cancer (NSCLC) is the largest subgroup of lung cancers, responsible for about 80% of all lung cancers. Targeted cancer therapy has opened a new window into treatment of non - small cell lung cancer. This therapy is very effective for patients who have EGFR (Epidermal growth factor receptor) activating mutations, primarily located in the tyrosine kinase domain in the form of a base - pair deletion at exon 19 (LREA deletions, accounting for about 54%) or a point mutation at exon 21 (L858R, accounting for about 43%). EGFR activating mutations occur in about 20% of NSCLC patients.

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Corresponding author: Tran Huy Thinh, Dept of

Biochem-istry, Center for Gene and Protein Research, Hanoi

Medi-cal University

E-mail: tranhuythinh@hmu.edu.vn

Received: 02 November 2016

Accepted: 10 December 2016

TARGETING TKI RESISTANCE IN NON - SMALL CELL

LUNG CANCER PATIENTS CAUSED BY SECONDARY EGFR

T790M MUTATION

Tran Quoc Dat, Tran Huy Thinh, Le Hoan,

Tran Van Khanh, Ta Thanh Van

Hanoi Medical University Non - small cell lung cancer (NSCLC) is the largest subgroup of lung cancers, responsible for about 80% of all lung cancers Targeted cancer therapy has opened a new window into treatment of non - small cell lung cancer This therapy is very effective for patients who have EGFR (Epidermal growth factor receptor) activat-ing mutations, primarily located in the tyrosine kinase domain in the form of a base - pair deletion at exon 19 (LREA deletions, accounting for about 54%) or a point mutation at exon 21 (L858R, accounting for about 43%) EGFR activating mutations occur in about 20% of NSCLC patients However, acquired resistance to EGFR tyrosine kinase inhibitors (TKIs) in NSCLC patients prevails after 10 - 20 months of treatment, in rela-tion to several genetic alterarela-tions of the tumors The most frequent genetic alterarela-tion is the secondary mutation T790M in exon 20 of the EGFR gene In this study, we detected secondary T790M resistance mutations in 11 EGFR - mutant patients who had poor response to targeted therapy We also report two typical clinical cases with TKI acquired resistance, using a standard sequencing - based method followed by the Scorpions ARMS (Scorpions - Amplification Refractory Mutation System) method, which has enhanced analytical sensitivity The T790M mutation analysis of specimens from 21 patients detected a total of 10 mutants (47.6%), including 4 mutants that were detected by both sequencing and the Scorpions ARMS method and 6 mutants that were only detected by the Scorpions ARMS method.

Keywords: Non - small cell lung cancer, TKI - acquired resistance, Secondary T790M mutation

I INTRODUCTION

Lung cancer is the leading cause of cancer

mortality worldwide In the United States, lung

cancer accounted for an estimated 159,480

deaths in 2013, which is more than breast,

colon, prostate and pancreatic cancer deaths

combined [Cancer Fact & Figure 2013].

Conventional treatment for advanced NSCLC

has typically consisted of chemotherapy [1]

However, despite proven benefits in an appro-priately selected population, the impact of tra-ditional chemotherapy on progression - free survival (PFS) and overall survival (OS) remains small Currently, the 1 - year survival rate for a patient with stage IIB/IV NSCLC is,

at best, around 30%, with a median survival time of no more than 12 months [2; 3] Recently, targeted cancer therapy has opened

a new window into treatment the of NSCLC This therapy is very effective for patients that have epidermal growth factor receptor (EGFR) activating mutations, the majority of which are located in the tyrosine kinase domains in the form of a base-pair deletion at exon 19 (LREA deletions, accounting for about 54% of EGFR

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activating mutations) or a point mutation at

exon 21 (L858R, accounting for about 43% of

EGFR activating mutations) EGFR activating

mutations occur in about 20% of NSCLC

patients, with significantly increased proportions

in those patients with adenocarcinoma on

histology, female patients, patients of Asian

ethnicity, and patients who were never

-smokers [4; 5] Even though sensitizing

mutations in the EGFR gene are associated

with higher response rates to EGFR - TKI

therapies and prolonged progression - free

survival, resistant mutations eventually prevail

via cellular survival pressure [6]

Many studies have postulated various

possible resistance mechanisms, including: a

secondary mutation in the EGFR gene

(a T790M mutation in exon 20, a L747S

mutation, a D761Y mutation in exon 19, or a

T854A mutation in exon 21), amplification of

signaling molecules that bypass EGFR

inhibition (MET and HER2), mutations in other

genes that may substitute as oncogenic

drivers (PIK3CA and B - RAF), epithelial -

to-mesenchymal transition (EMT), and

conver-sion to small - cell lung cancer [7; 8] Among

them, the T790M point mutation in EGFR exon

20 is believed to be the main source of

resistance, accounting for over half of the

resistance seen to gefitinib and erlotinib The

aim of this study was to identify secondary

EGFR - T790M mutations in re - biopsied tissue

samples from Vietnamese NSCLC patients with

acquired resistance to TKis Responsible

molecular alterations and mechanisms used to

overcome TKI resistance were contemporarily

assessed We also report two clinical cases of

women with NSCLC whose histories showed a

II MATERIALS AND METHODS

1 Materials

Twenty - one re - biopsied formalin - fixed paraffin embedded (FFPE) tissue samples were obtained from NSCLC patients with acquired resistance to EGFR - targeted therapy The patients came from Huu Nghi Hospital, National Bach Mai Hospital, or the National Cancer Hospital All three hospitals are located in Vietnam Patients were diagnosed with NSCLC, adenocarcinoma stage IIIB - IV, based on clinical features and pathology results These patients had all been previously biopsied to identify EGFR sensitizing mutations (11/21 patients had LREA deletion mutations in EGFR exon 19, and 10/21 patients had L858R point mutations

in EGFR exon 21) All patients had been treated with EGFR - TkI, with their responses evaluated using the WHO/RECIST Criteria, and rebiopsied to identify the EGFR - T790M resistance mutation Jackman’s Evaluation Criteria was used to define more precisely the patients’ acquired resistance to EGFR - TkI (Table 1) [9]

2 Methods

DNA was extracted from biopsied FFPE samples using a Qiamp DNA Mini Kit (Qiagen, Hilden, Germany) according to the protocol for tissue samples in the manufacturer's instruc-tions The DNA obtained was eluted in 50 µL

of sterile bidistilled buffer, and the concentra-tion and purity of the extracted DNA were as-sessed by spectrophotometry The extracted DNA was stored at −20°C until use

Sequencing method: The EGFR gene was

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directly sequenced using an Advant 3100

automated sequencer (Applied Biosystems

Inc., Foster City, California, USA) Sequences

were aligned and inspected using a reference

sequence from GeneBank (NM_005228).

Scorpion ARMS (Scorpion Allele

Refractory Mutation System) method: We

used an EGFR Scorpion TM Kit (EGFR RGQ

PCR Kit, Qiagen, Germany), which combines

two technologies - ARMS and Scorpion - to

detect mutations in real - time PCR reactions,

with an analytical sensitivity of approximately

0.01% All reactions were carried out in 25 µL volumes with 1 µL of template DNA, 10 µL of primer mix and 0.1 µL of Taq polymerase Real - time PCR was carried out under the following conditions: initial denaturation at 95°

C for 10 minutes, 50 cycles of 95°C for 30 seconds and a final 60 seconds at 62°C with fluorescence reading (set to FAM dye (6 - carboxy fluorescein) that allows optical excitation at 480 nm and measurement at 520 nm) at the end of each cycle

Table 1 Jackman’s Evaluation Criteria to define acquired resistance to

EGFR - TKi in NSCLC

1 Previous treatment with a single - agent EGFR TKI (eg, gefitinib or erlotinib)

2 Either or both of the following:

+ A tumor that harbors an EGFR mutation known to be associated with drug sensitivity or objective clinical benefit from treatment with an EGFR TKI;

+ Systemic progression of disease (Response Evaluation Criteria in Solid Tumors [RECIST] or WHO) while on continuous treatment with gefitinib or erlotinib within the last 30 days

4 No intervening systemic therapy between cessation of gefitinib or erlotinib and initiation of new therapy

III RESULTS

Twenty - one patients with NSCLC with

acquired resistance to EGFR - targeted

ther-apy were enrolled in this study Rates of

detection of the T790M mutation in exon 20 of

the EGFR gene using the sequencing method

and the Scorpion ARMS method are shown in

table 2

Our results showed that T790M mutation analysis of the specimens from the 21 patients

in our study detected 10 mutants (47.6%), including 4 mutants that were detected by both sequencing and the Scorpions ARMS method and 6 mutants that were only detected by the Scorpions ARMS method because of the limited cancer tissues obtained after re -biopsy

3 Research Ethics: This research was approved by the ethics committee of the Hanoi

Medical University, decree No.161/HMUIRB, signed on February 15th, 2014

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Table 2 Patient subtype pathology and EGFR mutation status before

and after treated with TKis

pathology

Stage

Mutation identi-fied before treated with EGFR- TKis

Mutation identified after treated with

EGFR- TKis Sequencing

method

Scorpions ARMS method

1 Adenocarcinoma IIIB LREA LREA LREA, T790M

2 Adenocarcinoma IV LREA LREA LREA

3 Adenocarcinoma IV L858R L858R L858R

4 Adenocarcinoma IV LREA LREA LREA, T790M

5 Adenocarcinoma IIIB L858R L858R, T790M L858R, T790M

6 Adenocarcinoma IV LREA LREA LREA

7 Adenocarcinoma IV L858R L858R L858R, T790M

8 Adenocarcinoma IV L858R L858R L858R

9 Adenocarcinoma IIIB LREA LREA, T790M LREA, T790M

10 Adenocarcinoma IIIB LREA LREA LREA

11 Adenocarcinoma IV L858R L858R L858R

12 Adenocarcinoma IIIB LREA LREA LREA

13 Adenocarcinoma IV L858R L858R L858R

14 Adenocarcinoma IIIB LREA LREA LREA, T790M

15 Adenocarcinoma IIIB LREA LREA LREA, T790M

16 Adenocarcinoma IV L858R L858R L858R

17 Adenocarcinoma IIIB L858R L858R L858R

18 Adenocarcinoma IV L858R L858R, T790M L858R, T790M

19 Adenocarcinoma IV LREA LREA LREA

20 Adenocarcinoma IIIB L858R L858R L858R, T790M

21 Adenocarcinoma IV LREA LREA, T790M LREA, T790M

Type of mutation: LREA (E746_A750del): in - frame deletion mutation in exon 19; L858R (c.2573T > G): point mutation in exon 21; T790M (c.2369C > T): point mutation in exon 20 Reference sequence: (NM_005228)

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Figure 1 Results of the direct sequencing method and the Scorpion Amplified Refractory Mutation System (ARMS) method in patient no.05 before (A) and after (B) treated with

EGFR-Tkis

Before treatment with EGFR - TKis, a sensitizing mutation (L858R exon 21) was detected in this patient’s tumor After treatment with EGFR - TKis, not only was the L858R sensitizing mutation seen, but an EGFR - T790M secondary mutation was also detected in the re - biopsied tissue sample These mutations were seen using both the sequencing method and the Scorpion ARMS method This mutation is associated with resistance to EGFR - TKis.

Figure 2 Results of the direct sequencing method and the Scorpion Amplified Refractory Mutation System (ARMS) method in patient no.01 before (A) and after (B) treated with

EGFR-TKis

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Before treatment with EGFR - TKis, a deletion mutation in EGFR exon 19 was detected in the patient’s biopsied tissue This activating mutation is predicted to be a sensitizing mutation to EGFR-TKI 12 months after treatment with the EGFR TKI, when tissue was re - biopsied, sequencing revealed a suspicious signal in amino acid 790 in exon 20 This result was verified using the Scorpion ARMS method and the results showed two mutations in the re - biopsied tissue: the deletion mutation in exon 19 and a T790M secondary mutation.

Patient no.05 is one of the most typical

clinical cases of a NSCLC patient with

acquired resistance to TKIs caused by a

secondary EGFR - T790M mutation At the

time of diagnosis, the patient had symptoms of

persistent sinus congestion, cough, and mild

progressive dyspnea Computed tomography

(CT) of the chest revealed a large left upper

lobe pulmonary mass with bilateral pulmonary

metastases and right hilar, prevascular,

pretracheal, and subcarinal lymphadenopathy

CEA concentration was found to be 27.7 ng/

mL, and no distant metastatic lesions were

detected CT - guided biopsy of the primary

mass revealed a poorly differentiated

non - small - cell carcinoma, consistent with an

adenocarcinoma Point mutation L858R in

EGFR exon 21 was detected in the biopsy

tissue from this patient This EGFR activating

mutation is believed to be a sensitizing

mutation to EGFR tyrosine kinase inhibitors

The patient was started on oral Gefitinib for first - line treatment (250mg daily)

After 6 months, the patient had a complete response The symptoms of persistent sinus congestion, cough and dyspnea decreased However, after 13 month of gefitinib treatment, the patient developed acquired resistance to TKIs A lesion appeared in the left lower lobe, along with left pleural effusion and a mediastinal lymph node metastasis near the right lung Re - biopsy of the second lesion located in left lower lobe revealed adenocarcinoma The re - biopsied tissue was found to have a T790M mutation in exon 20, which is associated with resistance to first-generation EGFR ‑ TKIs The re - biopsied tissue was also found to have an activating mutation (L858R exon 21) These two mutations were detected by both the sequen-cing method and the Scorpion ARMS method

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Figure 3 Chest computed tomography (CT) scan of patient no.05

(A) before treatment with EGFR - TKis, (B) 6 months after starting gefitinib, the primary tumor decreased in size (C) 13 months after treatment with gefitinib, disease progression was seen

C

Figure 4 Chest computed tomography (CT) scan and bone scintigraphy of patient no.01

(A) before treatment with EGFR - TKis, (B) 6 months after starting gefitinib, the tumor decreased in size (C) 15 months after treatment with erlotinib, disease progression was seen.

Patient no.01 was a 38 year old woman

with no notable occupational exposure, no

family history of lung cancer, and no history of

smoking or exposure to second - hand smoke

The patient visited the National Cancer

Hospi-tal for evaluation of a chronic cough The initial

PET-CT scan showed a 3,5 x 5 cm tumor in

the left middle lobe that was connected to the pleura Left pleural effusion was also seen No affected lymph nodes were detected How-ever, the tumor had already spread to the bone and metastases were detected in the spine (fig 4) The patient was diagnosed with lung adeno-carcinoma by transbronchial lung biopsy and

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an activating EGFR mutation (exon 19

dele-tion) was detected using both the

sequencing method and the Scorpion ARMS

method The patient was started on Erlotinib

followed by Bisphotphonat for firstline

treatment, and her response to treatment was

evaluated using the RECIST Criteria

Progression - free survival (PFS) was

measured from the first day of target treatment

to the day of disease progression After 6

months, the patient had a complete response

and stable disease At the time of the

15 monthS follow-up after treatment with

er-lotinib was begun, the patient began to

ex-perience severe thoracic pain, shortness of

breath, and analgesia refractory to medication

A PET - CT scan revealed new metastatic

lesions in the contralateral lung and spine (fig

4C) A new biopsy was taken Besides the

previously identified activating mutation in

exon 19 of the EGFR gene, an additional

secondary T790M point mutation in exon 20 was

now seen, conferring resistance to erlotinib In

this case, the secondary T790M mutation was

confirmed only by the Scorpion ARMS method

with an analytical sensitivity of approximately

0.01% This was because of the limited cancer

tissues obtained after re - biopsy

IV DISCUSSION

Many previous studies have found that a

secondary mutation in the EGFR gene

(T790M mutation) and amplification of the

MET proto - oncogene could be the causes of

resistance mechanisms in small cell lung

cancer transformation Moreover, the T790M

point mutation in exon 20 of the EGFR gene is

the main mutation believed to cause

tumor cells’ acquired resistance to TKIs: i) the T790M mutation can change the kinase domain’s interactive region and block the binding of the EGFR TKI, so the transduction pathway can still be activated and cancer cells can continue with proliferation, migration, invasion and angiogenesis; ii) changing amino acid 790 from tyrosine to methionine physically impedes access to the ATP binding site by TKI drugs such as Erlotinib and Gefitinib [12] In this study, both the sequencing method and the Scorpion ARMS method was used to look for mutations However, the detection rate of the T790M mutation was lower than that found in previous studies, possibly due to sample size limitations [11; 13] Overall, our results suggest that the scorpion ARMS method should be used to detect secondary T790M -EGFR mutations instead of the direct sequencing method The direct sequencing method can be used initially as a screening method to detect EGFR mutations

V CONCLUSION

In this study, secondary EGFR-T790M mutation analysis of specimens from

21 Vietnamese NSCLC patients detected 10 mutants (47.6%), including 4 mutants that were detected by both sequencing and the Scorpions ARMS method and 6 mutants that were only detected by the Scorpions ARMS method because of the limited cancer tissues obtained after re - biopsy Further studies should be conducted better elucidate rates of EGFR mutations in NSCLC patients in Vietnam

Acknowledgements

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their voluntary involvement in this study This

work was supported by Research Grant

KC.04.16/11-15 from the Ministry of Science

and Technology in Vietnam

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