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Conservation tillage and residue management towards low greenhouse gas emission; storage and turnover of natural organic matter in soil under sub-tropical ecosystems: A review

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Soil organic carbon (SOC) dynamics in croplands is a crucial component of global carbon (C) cycle. Depending on local environmental conditions and management practices, typical C input is generally required to reduce or reverse C loss in agricultural soils. Changes in the soil organic carbon (SOC) stock are determined by the balance between the carbon input from organic materials and the output from the decomposition of soil C. The fate of SOC in cropland soils plays a significant role in both sustainable agricultural production and climate change mitigation. Tillage systems can influence C sequestration by changing aggregate formation and C distribution within the aggregate. Results showed that the soil organic carbon (SOC) stock in bulk soil was 40.2-51.1% higher in the 0.00-0.05 m layer and 11.3-17.0% lower in the 0.05-0.20 m layer in NT system no-tillage without straw (NT-S) and with straw (NT+S), compared to the MP system moldboard plow without straw (MP-S) and with straw (MP+S), respectively. Residue incorporation caused a significant increment of 15.65% in total water stable aggregates in surface soil (0– 15 cm) and 7.53% in sub-surface soil (15–30 cm).

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Review Article https://doi.org/10.20546/ijcmas.2019.804.259

Conservation Tillage and Residue Management towards Low Greenhouse Gas Emission; Storage and Turnover of Natural Organic Matter in Soil

under Sub-tropical Ecosystems: A Review

S.K Tomar 1* , N.C Mahajan 2 , S.N Singh 3 , Vinay Kumar 4 and R.K Naresh 5

1

KVK Belipar, Gorakhpur, 3 KVK Basti, 4 KVK Akbarpur, Narendra Dev University of

Agriculture & Technology, Kumarganj, Ayodhya, U.P., India 2

Institute of Agricultural Sciences; Department of Agronomy, Banaras Hindu University,

Varanasi-(U.P), India 5

Department of Agronomy, Sardar Vallabhbhai Patel University of Agriculture & Technology,

Meerut, (UP), India

*Corresponding author

A B S T R A C T

International Journal of Current Microbiology and Applied Sciences

ISSN: 2319-7706 Volume 8 Number 04 (2019)

Journal homepage: http://www.ijcmas.com

Soil organic carbon (SOC) dynamics in croplands is a crucial component of global carbon (C) cycle Depending on local environmental conditions and management practices, typical C input is generally required to reduce or reverse C loss in agricultural soils Changes in the soil organic carbon (SOC) stock are determined by the balance between the carbon input from organic materials and the output from the decomposition of soil C The fate of SOC in cropland soils plays a significant role in both sustainable agricultural production and climate change mitigation Tillage systems can influence C sequestration by changing aggregate formation and C distribution within the aggregate Results showed that the soil organic carbon (SOC) stock in bulk soil was 40.2-51.1% higher in the 0.00-0.05 m layer and 11.3-17.0% lower in the 0.05-0.20 m layer in NT system no-tillage without straw (NT-S) and with straw (NT+S), compared to the MP system moldboard plow without straw (MP-S) and with straw (MP+S), respectively Residue incorporation caused a significant increment of 15.65% in total water stable aggregates in surface soil (0–

15 cm) and 7.53% in sub-surface soil (15–30 cm) In surface soil, the maximum (19.2%) and minimum (8.9%) proportion of total aggregated carbon was retained with >2 mm and 0.1–0.05 mm size fractions, respectively DSR combined with zero tillage in wheat along with residue retention (T 6 ) had the highest capability to hold the organic carbon in surface (11.57 g kg -1

soil with the highest stratification ratio of SOC (1.5) A considerable proportion of the total SOC was found to be captured by the macro-aggregates (>2–0.25 mm) under both surface (67.1%) and sub-surface layers (66.7%) leaving rest amount in micro- aggregates and ‗silt + clay‘ sized particles Soil tillage practices have a profound influence on the greenhouse gas (GHG) balance However there have been very few integrated studies on the emission of carbon dioxide (CO 2 ), methane (CH 4 ) and nitrous oxide (N 2 O) and soil biophysical and chemical characteristics under different soil management systems Tillage played a significant role in the flux of

CO 2 and CH 4 In contrast, N 2 O flux was determined mainly by microbial biomass carbon and soil moisture content Compared with other treatments, NT significantly reduced CH 4 emission among the rice growing seasons However, much higher variations in N 2 O emission were observed across the rice growing seasons due to the vulnerability of N 2 O to external influences The amount of CH 4 emission in paddy fields was much higher relative to N 2 O emission Conversion of CT to NT significantly reduced the cumulative

CH 4 emission for both rice seasons compared with other treatments The mixing of residues/surface retention into the soil increases SOM mineralisation due to greater exposure to microbial decomposers and optimal moisture and temperature regimes Soil disturbance by tillage leads to destruction of the protective soil aggregate This in turn exposes the labile C occluded in these aggregates to microbial breakdown The present study found that SOC change was significantly influenced by the crop residue retention rate and the edaphic variable of initial SOC content

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Introduction

Agriculture accounts for approximately 40-50

% of the earth‘s surface is managed for

agricultural purposes and contributes 10-12 %

of global greenhouse gas (GHG) emissions,

around 5.1-6.1 Pg CO2 -eq yr-1 in 2005 (Smith

et al., 2007a) This is made up of 3.3 Pg CO2

-eq yr-1 from methane (CH4) and 2.8Pg CO2

-eq yr-1 from nitrous oxide (N2O) emissions

Although there are large exchanges of carbon

dioxide (CO2) between the atmosphere and

agricultural ecosystems, emissions are

thought to be roughly balanced by uptake,

giving a net flux of only around 0.04 Pg CO2

yr-1, less than 1 % of global anthropogenic

CO2 emissions (Smith et al., 2007a) Land use

change is accounted for separately, but

change to cultivated land is thought to

contribute a further 5.9 ± 2.9PgCO2-eq yr-1,

6-17 % of total global GHG emissions (Bellarby

et al., 2008) If indirect emissions from

agrochemical production and distribution and

on-farm operations, including irrigation, are

also included, an extra 0.4-1.6 Pg CO2-eq yr-1

(0.8-3.2 %) can be attributed to agriculture,

meaning that, in total, direct and indirect

emissions from agricultural activity and land

use change to agricultural use could

contribute as much as 32.2 % of all GHG

emissions (Bellarby et al., 2008) Agriculture

is the main source of global non CO2 GHG

emissions, contributing around 47 % of

anthropogenic CH4 emissions and 58 % of

N2O, although there is a large degree of

uncertainty around estimates for both

agricultural contribution and total

anthropogenic emissions The main sources,

N2O from soils and CH4 from enteric

fermentation, make up around 70 % of

non-CO2 emissions from the sector, with biomass

burning, rice cultivation, and manure

management, accounting for the remainder

(Smith et al., 2007a) Conservation tillage is

one among many different mitigation options

suggested to reduce GHG emissions from

agriculture Conservation tillage practices such as reduced/minimum/zero tillage, direct drilling and strip cropping are also widely recommended to protect soil against erosion

and degradation of structure (Petersen et al.,

2011), create greater aggregate stability

(Fernandez et al., 2010; Zotarelli et al., 2007)

increase soil organic matter content, enhance

sequestration of carbon (Six et al., 2000) mitigate GHG emissions (Kong et al., 2009) and improve biological activity (Helgason et

al., 2010)

Minimum tillage practices have been reported

to reduce GHG emissions through decreased use of fossil fuels in field preparation and by increasing carbon sequestration in soil

(Petersen et al., 2008) The crop residues

accumulated on the soil surface under reduced tilled conditions may result in carbon being lost to the atmosphere upon decomposition

(Petersen et al., 2008) Furthermore, climate

change mitigation benefits such as reduced

CO2 emissions, by virtue of increased sequestration of carbon and increased CH4

uptake under reduced tillage, could be offset

by increased emissions of N2O, a greenhouse gas with higher warming potential than both

CO2 and CH4 (Hermle et al., 2008; Chatskikh

and Olesen, 2007) Increased N2O emissions have been linked to increased denitrification under reduced tillage due to the formation of micro-aggregates within macro-aggregates

that create anaerobic micro sites (Hermle et

al., 2008) with increased microbial activity

leading to greater competition for oxygen (West and Marland, 2002)

Reduction of tillage can also create increased soil densification and a subsequent decrease

in the volume of macro-pores (Schjønning and Rasmussen, 2000) leading to reduction in gaseous exchange Soil aggregation and the resultant geometry of the pore structure are vitally important characteristics affected by tillage practices which impact on the physico-

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chemical and hydro-thermal regime in soil,

and ultimately crop yield Additionally, the

effect of tillage on the environment varies

across farms geographically since the impacts

of cultivation on soil organic matter and net

greenhouse balance depends on soil type,

climatic variables and management

(Chatskikh and Olesen, 2007)

Natural organic matter in soils is the largest

carbon reservoir in rapid exchange with

atmospheric CO2, and is thus important as a

potential source and sink of greenhouse gases

over time scales of human concern (Fischlin

and Gyalistras, 1997) SOM is also an

important human resource under active

management in agricultural and range lands

worldwide Questions driving present

research on the soil C cycle include: Are soils

now acting as a net source or sink of carbon

to the atmosphere? What role will soils play

as a natural modulator or amplifier of climatic

warming? How is C stabilized and

sequestered, and what are effective

management techniques to foster these

processes? Answering these questions will

require a mechanistic understanding of how

and where C is stored in soils SOM quantity

and composition reflect the long-term balance

between plant carbon inputs and microbial

decomposition The processes underlying soil

carbon storage and turnover are complex and

dynamic, involving influences from global to

molecular scales At the broadest level, SOM

cycling is influenced by factors such as

climate and parent material, which affect

plant productivity and soil development At a

more proximate level, factors such as plant

species and soil mineralogy affect

decomposition pathways and stabilization

processes The molecular characteristics of

SOM play a fundamental role in all processes

of its storage and stability

Historical global estimates for the top meter

of soil vary from 800 Pg C to 2,400 Pg C,

converging on the range of 1,300–1,600 Pg C

to 1 m Batjes (1996) estimated that an additional 900 Pg C is stored between 1 and 2

m depth, and Jobbágy and Jackson (2000) revised that estimate to 500 Pg between 1 and

2 m and another 350 Pg between 2 and 3 m depth Global organic carbon stocks to 3 m are currently estimated at 2,300 Pg, with an additional 1,000 Pg contained in permafrost and peat lands (Jobbagy and Jackson, 2000;

Zimov et al., 2006) In this review paper we

sought to evaluate the impact of conservation tillage on storage and turnover of natural organic matter in soil and GHG emissions

We hypothesized that conservation tillage improves storage and turnover of natural organic matter in soil and reduces GHG emissions compared with conventional tillage through the enhanced development of the soil carbon associated with less anthropogenic disturbance

Reicosky and Archer (2007) reported that the

CO2 released immediately following tillage increased with ploughing depth and in every case was substantially greater than that from the no-tillage treatment Intensive soil cultivation breaks down soil organic matter (SOM), producing CO2, and consequently reduces the total C content There are many reports suggesting that soil tillage accelerates organic C oxidation, releasing large amounts

of CO2 to the atmosphere over a few weeks

(La Scala et al., 2008) Conservation tillage

has been shown to result in a greater percentage of soil present in macro-aggregates and a larger proportion of carbon associated with micro-aggregates compared to

that in conventional ploughing (He et al.,

2011) Under conventional ploughing, aggregates are readily broken down prior to micro-aggregate formation This leads to a reduction in the proportion of C that is more protected in micro-aggregates and thus to the

macro-loss of recalcitrant SOC (Chivenge et al., 2007) Li et al., (2011) investigated methane

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emission patterns in a double-rice cropping

system under conventional tillage and

no-tillage in south-east China, where no-no-tillage

reduced seasonal methane fluxes by 29% and

68% for the early and late rice, respectively

Ahmad et al., (2009) also found that

no-tillage significantly reduced methane

emissions from paddy fields compared to

conventional tillage (Fig 1, 2 and 3)

Sarkhot et al., (2012) reported that the

prepared nutrient enriched bio-char by

shaking the bio-char with dairy manure

effluent for 24 h, which increased the C and N

content of the bio-char by 9.3% and 8.3,

respectively When the untreated bio-cha and

N enriched bio-char were added to a soil in

eight week incubation, the reduction in

availableNH4+-N and NO3 —

N content was observed, suggesting the possibility of N

immobilization Still, N enriched bio-char

could be used as a slow release N fertilizer

The net N nitrification rates in the CK, 1%

BC and 3% BC treatments also peaked at day

25, then dramatically decreased and stayed at

a very low level (0.35–0.42mg/(kg d)) at the

end of incubation

Sander et al., (2014) reported that

incorporation of rice residues immediately

after harvest and subsequent aerobic

decomposition of the residues before soil

flooding for the next crop reduced CH4

emissions by 2.5–5 times and also improved

nutrient cycling in paddy field It was also

reported that residue incorporation

accelerated CH4 and N2O emissions from

irrigated rice field compared to residues left

on the soil surface The open burning of crop

residues emits CO2, CH4, and N2O

Mangalassery et al., (2014) also found that

neither ammonium (NH4-N) nor nitrate (NO3

-N) content in soil was affected by tillage Soil

from the upper 10 cm contained significantly

higher NH4-N than the 10–20 cm layer

Nitrate (NO3-N) followed a similar trend to

NH4-N Tillage type and duration did not influence the NO3-N content Soil depth significantly influenced NO3-N content with highest amount in the surface layer (0–10 cm) under both zero tillage and conventional tillage Considering the GHGs together, tilled soil produced 20% greater net global warming than zero tilled soil indicating a potential for zero tillage system to mitigate climate change after only 5 to 10 years since conversion Del

Grosso et al., (2005) also reported a 33%

reduction in global warming potential under zero tillage (0.29 MgCha-1yr-1) compared with tilled soil (0.43 Mg C ha-1 yr-1) for major non-rice cropping systems Also in sub-tropical conditions, zero tillage has been found to

reduce GWP by c 20% (Pivea et al., 2012)

Residues management and crop rotations can affect N2O emissions by altering the availability of NO3 −

in the soil, the decomposability of C substrates (Firestone and Davidson, 1989) The reduction of N2O to

N2 is inhibited when NO3 −

and labile C

concentrations are high (Senbayram et al.,

2012) The retention of crop residues and higher soil C in surface soils with CA play major roles in these processes Under anaerobic conditions associated with soil water saturation, high contents of soluble carbon or readily decomposable organic matter can significantly boost de-nitrification

(Dalal et al., 2003) with the production of

N2O favoured with high quality C inputs (Bremner, 1997) The quantity and quality of residues or cover crops of CA systems can also affect N2O emissions Legume residues can result in higher N2O–N losses (Millar et

al., 2004) than those from non-legume, low N

residues (Aulakh et al., 2001) Crop residues

may affect CH4 oxidation in upland soils and emission patterns in flooded soils differently depending on their C/N ratio; residues with a high C/N ratio have little effect on oxidation while residues with a narrow C/N ratio seem

to inhibit oxidation (Hiitsch, 2011) Grace et

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al., (2012) estimated an average of 29.3 Mg

ha−1 of GHGs emitted over 20 years in

conventional rice-wheat systems across the

IGP; this decreased by only 3% with the

widespread implementation of CA

Agricultural practices such as tillage and

fertilization have to be considered Food

systems alone – everything from growing

plants to the disposal of biomass – contribute

to 19–29% of global anthropogenic GHG

emissions Of this, 80–86% relate to

agricultural production (including indirect

emissions associated with land-cover change),

albeit with significant regional variation

(Vermeulen et al., 2012) On agricultural

sites, N2O emissions from legume-N were

significantly lower than fertilizer-N derived

N2O emissions (Schwenke et al., 2015)

Gupta et al., (2016) revealed that the GWP

(CH4 + N2O) of wheat–rice systems varied

from 944 to 1891 kg CO2 eq ha-1 and 1167–

2233 kg CO2 eq ha-1 in the first and second

years of wheat–rice cropping respectively

The combination of ZTW followed by DSR

showed significantly low GWP than other

combination of wheat and rice treatments

These combinations led to about 44–47%

reductions in GWP over the conventional

CTW-TPR system in both the years The

order of GWP among the different

combination of treatments was as follows:

(ZTW + RR) - DSR < DSR <

ZTW-IWD < ZTW + NOCUTPR + NOCU <

CTWTPR < ZTW-TPR in both the years The

share of rice in total GWP was 72–81% in

those combinations in which TPR was a

treatment while it varied from 56 to 65%

where DSR was a treatment These results

indicate that adoption of ZTW followed by

DSR in the IGP in place of conventional

CTW-TPR can be an efficient low carbon

emitting option With the development of new

drills, which are able to cut through crop

residue, for zero-tillage crop planting, burning

of straw can be avoided, which amounts to as

much as 10 tons per hectare, potentially reducing release of some 13–14 tons of

carbon dioxide (Gupta et al., 2004)

Elimination of burning on just 5 million hectares would reduce the huge flux of yearly

CO2 emissions by 43.3 million tons (including 0.8 million ton CO2 produced upon burning of fossil fuel in tillage) Zero-tillage on an average saves about 60 l of fuel per hectare thus reducing emission of CO2 by 156 kg per

hectare per year (Grace et al., 2003; Gupta et

al., 2004) Sah et al., (2014) revealed that the

CO2 emissions conventionally tilled (CT) wheat emitted the highest amount of CO2 (224

kg ha-1) followed by PRB (146 kg ha-1) and the lowest from ZT (126 kg ha-1) The highest

CO2 emission through CT attributed to higher tractor usage on land preparation and more pumping time on irrigation However, ZT and PBP wheat emitted lower CO2 to the atmosphere by 43.7 % and 34.9 %, respectively, as compared to CT

Conservation tillage practices decreased the exposure of un-mineralized organic substances to the microbial processes, thus reducing SOM decomposition and

CO2 emission Apart from C, other greenhouse gases (GHGs) notably, nitrous oxide (N2O) and methane (NH4), have been reported to be influenced by tillage regimes (Steinbach and Alvarez, 2006) About 38% of the emissions to the atmosphere can be ascribed to nitrous oxide from soils (Bellarby

et al., 2008) while methane is considered as

the most potential greenhouse gas after carbon dioxide (IPCC, 2001) Significantly higher N2O emissions from ploughed than no-

tilled sites has been reported by Kessavalou et

al., (1998) The higher aeration in tilled soil

increases oxygen availability, possibly resulting in increased aerobic turnover in the soil and thus an increased potential for

gaseous emissions (Skiba et al., 2002) Seidel

et al., (2015) compared the ratio between

greenhouse gas emissions from inputs and

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crop output across organic and conventional

cropping systems and suggests that a legume

tilled management exhibited the best ratio

(59%) followed by manure tilled (63%),

manure no till (65%), legume no till (84%)

and conventional till (90%) as a per-cent of

the GHG emissions from conventional no till

management

Several of the agricultural and forestry GHG

mitigation options provide ancillary

co-benefits to the agricultural sector and to

society, making them somewhat unique in

their ability to address climate change

simultaneously with other pressing social and

environmental issues This has earned these

reductions the title of ―charismatic carbon

credits.‖ Increasing soil C also increases

available plant nutrients; considering the

nutrient supplying capacity of just N, P, S,

a1% increase in soil organic matter content

(equivalent to 21 Tons of CO2) would

translate to 75 lb N, 8 lb P and 8 lb of S per

acre (Rice et al., 2007)

CO2 in the atmosphere is in a constant state of flux among its repositories, or ―sinks‖; this is called the Carbon Cycle The movement, or

―flux,‖ of carbon between the atmosphere and the land and oceans sinks is dominated by natural processes, such as plant photosynthesis While these natural processes can absorb some of the net 6.3 billion metric tons of human-produced CO2 emissions emitted each year (about 2 billion metric tons are absorbed by the ocean and 1 billion by terrestrial systems, including soils), that leaves an estimated 3.2 billion metric tons that are added to the atmosphere annually The Earth‘s positive imbalance between emissions and absorption of GHG has resulted in the increased concentration of greenhouse gases in the atmosphere This causes global climate change

Turnover time and dynamics of soil organic matter

Cambardella and Elliott, (1994) reported that the turnover time of POC ranged from 5 to 20 years in cultivated grassland soils The reason might be that after cultivation of virgin black soils, soybean (C3 crop) residues provided an extra source of organic matter input in addition to corn-derived C (C4 crop) It might also be due to a certain amount of black C in

POC (Knicker et al., 2005) The mean

turnover time indicated faster turnover of SOC in coarse fraction than that in fine fraction We suggested that short-term NT did not significantly affect the turnover time of SOC The turnover time of SOC was even longer in MP plots because of the incorporation of returned crop residues into soils Thus, the short-term impact of no tillage was firstly shown in the coarse-size fractions (POC) The distribution of C3–C mainly in fine particles (silt plus clay) indicated that the turnover of SOC in coarse-size fraction was faster under tillage practices Regardless of residue type, mineralization of SOM

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increased up to from 50 to 90% due to

addition of low and high levels, respectively,

whereas residue addition was increased 3.6

times Therefore, the amount of primed CO2

decreased per unit of applied residue This

was also reported by Guenet et al., (2010) and

Xiao et al., (2015)

Zhu et al., (2015) revealed that the soil total

organic C (TOC) and labile organic C fraction

contents were higher under the straw return

treatments compared to the no straw return

treatment (0% S) at a 0–21 soil depth The

50% annual straw return rate (50% S) had

significantly higher soil TOC, dissolved

organic C(DOC), easily oxidizable C (EOC),

and microbial biomass C (MBC) contents

than the 0% S treatment at a 0–21 cm depth

All of the straw return treatments had a

significantly higher DOC content than the

0%S treatment at a 0–21 cm depth, except for

the 100% only rice straw return treatment

(100% RS) Wang et al., (2015) also found

that in the early paddy field, the average

values of the total SOC, LFOC, DOC and

MBC concentration in the top 40cm soil were

significantly higher in the straw application

plots than in the controls, by 7.2% 8.8%,

15.6%, and 128.6%, respectively Wright et

al., (2007) reported that in the 0-5 cm soil

depth, no-tillage increased

macro-aggregate-associated OC as compared to conventional

tillage Macro-aggregates accounted for

38-64, 48-66, and 54-71% of the total soil mass

in the 0-5, 5-10, and 10-20 cm soil depths,

respectively The corresponding proportions

of the silt+clay fraction were 3-7, 2-6, and

1-5%, respectively Proportions of

macro-aggregates were increased with reduction of

soil tillage frequency For the 0-5 cm soil

depth, treatments NT and 4T had significantly

higher mass proportions of macro-aggregates

(36 and 23%, respectively) than that of

treatment With additions of crop residues, the

amount of macro-aggregates increased in all

tillage treatments Conservation tillage

significantly increased SOC concentration of bulk soil in the 0−5 cm soil layer This increase in SOC concentration can be attributed to a combination of less soil disturbance and more residues returned to the

soil surface under conservation tillage (Du et

al., 2010; Dikgwatlhe et al., 2014) Alvarez et al., (2009) also found that NT increases SOC

and total N concentrations in the first centimetres of the soil profile because NT

maintains surface residues Vanden Bygaart et

al., (2003) observe that non-inversion tillage

physically protects part of the organic matter

in the top layer from mineralization by inclusion within macro-aggregates With conventional inversion tillage on the other hand, aggregates will be more thoroughly disrupted, assisting loss of organic matter

Mangalassery et al., (2014) revealed that zero

tilled soils contained significantly more soil organic matter (SOM) than tilled soils Soil from the 0–10 cm layer contained more SOM than soils from the 10–20 cm layers in both zero tilled (7.8 and 7.4% at 0–10 cm and 10–

20 cm respectively) and tilled soils (6.6% at 0–10 cm and 6.2% at 10–20 cm)

Temporal scales of soil C dynamics

Wang et al., (2016) also found that higher

amounts of C input can lead to higher soil C sink capacities On a global average, the total amounts of C input to soils are 1.7, 2.7 and3.7 MgC ha−1 under the crop residue retention rates of 30, 60 and 90 %, respectively Lal, (2004) reported that the rates of SOC sequestration in croplands range from 0.02 to 0.76 MgC ha−1 yr−1 when improved systems

of crop management are adopted However, it should be noted that the increased SOC sequestration rate that is contributed to by the increased C input can be limited at longer periods, as the SOC would eventually reach a

relatively stable threshold (Stewart et al.,

2007) On a global scale, the estimated efficiency of the conversion of C input to

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SOC is 14 %, which falls within the 10–18 %

range estimated by Campbell et al., (2000) It

should be noted that the conversion efficiency

varies across space and is highly dependent

on the local climatic and edaphic conditions

(Yu et al., 2012) Mangalassery et al., (2014)

observed that zero tilled soils contained

significantly more microbial biomass carbon

than tilled soils The mean microbial biomass

carbon under zero tilled soil was 517.0 mg kg

-1

soil compared with 418.7 mg kg-1 soil in

tilled soils Microbial biomass carbon was

significantly higher in the 0–10 cm layer (517

mg kg-1 soil) than the 10–20 cm layer (419

mg kg-1 soil) under zero tillage and

conventional tillage Moreover, tillage and

soil depth significantly influenced soil

microbial biomass nitrogen Zero tilled soils

contained higher microbial biomass nitrogen

(91.1 mg kg-1 soil) than tilled soil (70.0 mg

kg-1 soil) Surface layers (0–10 cm)

maintained more microbial biomass nitrogen

than sub surface layers (10–20 cm) under both

zero tilled soils and tilled soils

Fortuna et al., (2003a) found that addition of

organic nutrient sources like compost to the

soil for more than 6 years has the potential to

increase the pools of slow (10% increase) and

resistant (30% increase) C and the potential

pool of potentially mineralizable N West and

Post (2002) calculated that converting from

mouldboard plough to no-till sequestered an

additional 0.57±0.14 Mg Cha-1yr-1 of C and

complex crop rotations had the potential to

sequester an additional 20±12 gCm-2yr-1 of C

Seventeen ±15% of C applied in animal

amendments such as poultry manure becomes

part of soil organic matter (SOM) (Johnson et

al., 2009) Key management practices that

retain or return residues to the soil have been

shown to insulate and elevate soil

temperatures reducing the extremity and

frequency of freeze-thaw cycles leading to a

reduction in N2O emissions Soil C and N

dynamics are influenced to a greater degree

by quantity rather than quality of plant

residues Gentile et al., (2011) reported that

the quality of crop residues effects short term nutrient dynamics and has a less of an impact

on C sequestration Jha et al., (2012)

suggested that the addition of FYM to soil increased the active C pool to a greater extent

as compared to the slow and resistant C pools

Powlson et al., (2012) also found the effect of

reduced tillage and addition of different organic materials on soil C stocks and N2O emissions They found that reduced tillage practices increased the annual C stocks compared to conventional tillage However, this was compensated for increased N2O emissions under reduced tillage management

Dendooven et al., (2012) revealed that no till

with crop residue removal and conventional tillage with residue retention or removal were net sources of CO2, with a positive net GWP ranging from 1.288 to 1.885 Mg CO2

ha−1yr −1 Hence, no till when practiced with residue retention had higher N2O emissions but also increased the C storage to an extent that the systems had net negative GWP

Gattinger et al., (2012) concluded that the

SOC stocks and C sequestration rates were significantly higher in the zero net input organic farming systems as compared to non-organic cropping systems by 1.98 ±1.50 MgCha-1 and 0.07±0.08 MgCha-1yr-1(mean ± 85% confidence interval) respectively Palm

et al., (2014) reported that the combined

effect of types of crops, intensity of cropping, duration of the cropping systems, the amount

of inputs added to the systems in the form of residues and the tillage intensity along with soil properties like soil texture, temperature and moisture determines the overall soil C

and N turnover and storage Thomazini et al.,

(2015) reported that organic no till with leguminous intercropping and pre-plant compost application had the potential to immobilize C in microorganisms thereby promoting a positive C balance in the soil leading to a C sink and improved soil health

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Zhao et al., (2016) indicated that returning

corn straw to the soil along with mixing it

reduced the CO2 emissions and increased the

soil organic carbon content thereby improving

the composition of micro-aggregate better

than straw mulching Zhang et al., (2016)

indicated that the application of chemical

fertilizers plus manure could be a suitable

management for ensuring crop yield and

sustaining soil fertility but the ratio of

chemical fertilizers to manure should be

optimized to reduce C and N losses to the

environment

Tillage system influence on soil organic

carbon storage

Wang et al., (2018) reported that tillage

system change influenced SOC content, NT,

ST, and BT showed higher values of SOC

content and increased 8.34, 7.83, and

1.64 Mg·C·ha−1, respectively, compared with

CT Among the 3 changed tillage systems, NT

and ST showed a 12.5% and 11.6% increase

in SOC content then BT, respectively Tillage

system change influenced SOC stratification

ratio values, with higher value observed in BT

and NT compared CT but ST Therefore, in

loess soil, changing tillage system can

significantly improve SOC storage and

change profile distribution Naresh et al.,

(2018) reported that conservation tillage

practices significantly influenced the total soil

carbon (TC), Total inorganic carbon (TIC),

total soil organic carbon (SOC) and

oxidizable organic carbon (OC) content of the

surface (0–15 cm) soil Wide raised beds

transplanted rice and zero till wheat with

100% (T9) or with 50% residue management

(T8) showed significantly higher TC, SOC

content of 11.93 and 10.73 g kg-1,respectively

in T9 and 10.98 and 9.38 g kg-1, respectively

in T8 as compared to the other treatments

Irrespective of residue incorporation/

retention, wide raised beds with zero till

wheat enhanced 53.6%, 33.3%, 38.7% and

41.9% of TC, TIC, SOC and OC, respectively, in surface soil as compared to conventional tillage with transplanted rice cultivation Simultaneously, residue retention caused an increment of 6.4%, 7.4%, 8.7% and 10.6% in TC, TIC, SOC and OC, respectively over the treatments without residue management

Concerning the organic carbon storage, SOCs varied between 31.9 Mg·ha−1 and 25.8 Mg·ha−1 under NT, while, in tilled treatments, SOCs ranged between 28.8 Mg·ha−1 and 24.8 Mg·ha−1 These values were lower than those observed by

Fernández-Ugalde et al., (2009) who found,

in silty clay soil, a SOCs at 0–30 cm of 50.9 Mg·ha−1 after 7 years of no tillage, which was significantly higher than the 44.1 Mg·ha−1 under CT under wheat-barley

cropping system in semiarid area Hernanz et

al., (2009) also found, after 11 years under

NT, a SOCs of 37 Mg·ha−1 which was higher than 33.5 Mg·ha−1 under CT, using a wheat-

vetch (Vectoria sativa L.) rotation in silty soil

The lower SOCs values we observed can be explained by the fact that more time is needed before achieving the peak sequestration rate

under NT Xu et al., (2013) observed that the

SOC stocks in the 0–80 cm layer under NT was as high as 129.32 Mg C ha−1,significantly higher than those under PT and RT The order

of SOC stocks in the 0–80 cm soil layer was

NT > PT > RT, and the same order was observed for SCB; however, in the 0–20 cm soil layer, the RT treatment had a higher SOC stock than the PT treatment. Alemayehu et

al., (2016) also found that the carbon storage per hectare for the four soil textures at 0 to 15

cm depth were 68.4, 63.7, 38.1 and 31.3 tha-1for sandy loam, silt loam, loam and clay loam; respectively Sand and silt loams had nearly twice the organic carbon content than loam and clay loam soil The soil organic carbon content for tillage type at 0 to 15 cm was 8.6, 10.6, 11.8 and 19.8 g kg-1 for deep

Trang 10

tillage, minimum tillage, shallow tillage, and

zero tillage; respectively Among tillage types

soil organic carbon storage could be increased

by using the minimum and shallow tillage

SOC storage decreased with soil depth, with a

significant accumulation at 0-20cm depth

Zheng et al., (2018) reported that across

treatments, aggregate-associated C at a depth

of 0–10cm was higher in the NT and ST

treatments than in the MP and CT treatments

The advantage of the NT treatment weakened

with soil depth, while the amount of

aggregate-associated C remained higher for

the ST treatment There were more

macro-aggregates in the ST and NT treatments than

in the MP and CT treatments, while the MP

and CT treatments had more

micro-aggregates The sum of macro-aggregate

contributing rates for soil organic C (SOC)

was significantly superior to that of the

micro-aggregates Mahajan et al., (2019) reported

that the increased SOC stock in the surface 50

kg m-2 under ZT and PRB was compensated

by greater SOC stocks in the 50-200 and

200-400 kg m-2 interval under residue retained, but

SOC stocks under CT were consistently lower

in the surface 400 kg m-2.Soil organic carbon

fractions (SOC), microbial biomasses and

enzyme activities in the macro-aggregates are

more sensitive to conservation tillage (CT)

than in the micro-aggregates Responses of

macro-aggregates to straw return showed

positively linear with increasing SOC

concentration Straw-C input rate and clay

content significantly affected the response of

SOC

Soil organic carbon and sequestration

SOM is a complex mixture which contributes

positively to soil fertility, soil tilth, crop

production, and overall soil sustainability It

minimizes negative environmental impacts,

and thus improves soil quality (Farquharson

et al., 2003) (Fig 4a) Loveland and Webb

(2003) suggested that a major threshold is 2%

SOC (ca 3.4% SOM) in temperate regions, below which potentially serious decline in soil quality will occur Storage of SOC is a balance between C additions from non-harvested portions of crops and organic amendments, and C losses, primarily through organic matter decomposition and release of respired CO2 to the atmosphere Organic matter returned to the soil, directly from crop residues or indirectly as manure, consists of many different organic compounds Some of these are digested quickly by soil microorganisms The result of this is a rapid formation of microbial compounds and body structures, important in holding particles together to provide soil structure and to limit soil erosion, and the release of carbon dioxide back to the atmosphere through microbial

respiration (Kladivko 2001) Paustian et al.,

(1998) compared tillage systems, ranging in duration from 5 to 20 years, and estimated that NT resulted in an average soil C increase

of 285 g/m2, compared to conventional tillage

(CT) Liu et al., (2003) showed a significant

decline of total SOC that occurred in the first

5 years of cultivation where the average SOC loss per year was about 2300 kg/ha for the 0–

17 cm horizon The average annual SOC loss between 5- and 14-year cultivation was 950 kg/ha and between 14- and 50-year cultivation it was 290 kg/ha These data clearly showed a rapid reduction of SOC for the initial soil disturbance by cultivation and a relatively gradual loss later Compared with

organic matter in the uncultivated soil, Liu et

al., (2003) also indicated that the total SOC

loss was 17%, 28%, and 55% in the 5-, 14- and 50-year cultivation periods, respectively The latter would correspond to the release of approximately 380 ton CO2/ha to the atmosphere

Within the surface 7.5 cm, the no-till system possessed significantly more SOC (by 7.28 Mg/ha), particulate organic matter C (by4.98 Mg/ha), potentially mineralizable N (by 32.4

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