LIST OF MAMMALS FROM SOMALLLAND, ELLIOT 1897

no Field Columbian Museum —Zoology, Vol.. 112 Field Columbian Museum —Zoology, Vol.. ii6 Field Columbian Museum —Zoology, Vol.. Onlythe males of these species carry horns, about 2.50 inc

Publication 19.

MUSEUM EAST AFRICAN EXPEDITION TO SOMALI-LAND

IN 1896 Bv D G ELLIOT, F R S E.

Thu chief reason that induced the Field Columbian Museum to

lost if examples of these still living specieswere to be procured for

the Institution, to show to those who come afterus the creatures that

ample measurements of thespecimens obtained Unfortunately, on

that would undoubtedly have been obtained are omitted from the list.

Many more examplesof the different species could easily have been

FAM SUID^.

Phacochoerus africanus (Gmel)

The Wart-Hog was fret^uentlynutwitli tliroughout thecountry

traveledbyus. Thefirstoneseenwas at Mandera near thebaseof

no Field Columbian Museum —Zoology, Vol i.

seen nae and sought refuge in an ahnost impenetrable thicket of

meninto the bushes from the otherdrovehimout. Hepresented

a grand appearance as he came,trottingmajestically along,

with-out the slightest evidence of fear, his tail held upright with the

tassel pendentforward, and his mane, very long and thick, raised

stood, and only the click of the hammer answered the pull of

shikari had forgotten to throw a shell into the chamber before

among the bushes, and we were obliged to track him a long way

to him, champing his tushes until the foam flew from his mouth,

as he made frantic but unavailing efforts to reach me Another

bullet laid him lifeless.

water, for we met them in the middle of the Haud where

cer-• tainly the nearest water-hole must have been 50 miles away, too

a dangerous antagonist, as he could inflict fearful injuries with

of Eastern Africa It obtains its trivial name from the peculiarwart-like excrescences upon various parts of the head These

peculiareffect, not however one of beaut3^ The nativesbeing

Mussulmanswill not touch a hog, oranything that has come in

these animals, either as a trophy or for scientific purposes, must

do all the work himself, while his followers squat around him,

generally Whenthemembers of alitterarrive atan agebetween

theyleave theparentsand themselves

At least this I imagine to be the case, for young pigs that I saw,

andsomeofwhichI killed,roamingabout without anyadults near,

number of young usually found in a litter, but I should supposefromfour to eightwouldbe nearly correct Many fall victims to

that the species is kept from undue increase

This species is evidently the PhacocJuTnis cvliani, Riipp, Atlas

I, pi. 25.=Sus africanus, Gm. Syst Nat., 1788, p. 220, havingtwo upper and four lowerincisors. There is, however, a curious

incisors well exhibited in both jaws, as has also the smallest

specimenonlya fewmonthsold. But of thethree otherexamples

over ayear old, two exhibit the lower incisors just emerging, but

no indication of any in the upper jaw, while the third gives no

indicationof anyin either jaw As this last and one of theothers

were killed out of the same sounder, the idea of two species cannot be for a moment entertained But it would seem to show

which three views are given, is, in its superior outline, very high

at the occiput, and descends rapidly to the posterior end of thenasals, these possessing a gradual curve throughout theirlength

INXHES

" " ' at anteriorrim of orbits (outline of molars) 7.80

Extentofupperpremolars i 40

Lengthofuppertushes betweentips 10.00

" " " " extreme outsidemeasurement 10.65

112 Field Columbian Museum —Zoology, Vol i.

FAM BOVID^.

This rare antelope, to obtain which I made a special trip to

Toyo Plain, is only found in a few localities on the elevated

to its great swiftness to carry it beyond the reach of its enemies

any place, so that stalking, inthe usual acceptation of theterm,

is practically an impossibility, as the Hartebeest see the hunter

Gen-erally they are first sighted along the horizon line of the plain,

theirpeculiar shape It is an ungracefulcreatureboth in

appear-ance and in its action when in motion The hind quarters are

canter, it possesses the greatest staying poAverand is really theswiftest of all the antelopes, always graduating itspace to that of

them, as the courage of the entire number is only equal to that

of the most timid, and as soon as onebegins to run all are off at

once Whenone is seen in the distance on the plain, the hunterwalks toward them in a direct line if theyare busy feeding ormoving slowly away, but should they stop to look around them,

to try a shot,they beginto run, in their rather slow,

clumsy-look-ing canter, but which is not usually kept up for anygreat

and it usually does many times before the quarry is secured, or

runs away altogether, the hunter has all his work to do over

again It is therefore only by the exercise of much patienceand

and it is practicallyuseless to follow such a one as it rarely

it inhabits there is no water, save what may occasionally fall inshowersduring the brief so-called rainyseason, and this antelope

must go practically without drinking, at all events for long

periods Those we killed were in excellent condition, but like

Swayne's Hartebeest can not be considered a game animal; there

is very little sport or excitement in its pursuit, and if I had not

4 and 5 feet high at the withers Both sexes possess horns,

and these vary according to age, both in the extent and manner

ofcurvature Some horns of bulls measure over23 inches from

and on account of the absence of fat is only fairly palatable

Madoqua swaynei (Thos.)

Madoqua swaynei Swayne's Dik-Dik Native name SakdroGiiyu.

Of the three species of pigmyantelopes inhabiting Somali-land

have no specimen from that land, whereitseemstobesupplanted

114 Field Columbian Museu.m—Zoology, \'<)l i.

in that direction it extends its range to Ogaden also It is

clearly distinguishable from M. pliillipsi, the color of its coat

quite large enough to make this fact conspicuously plain, and,

indepen-dent species and not races one of the other This

accompan-ied by a young one They would bound out of a clump of aloes

or from beneath some bush and go rushing away just likerabbits,

and it was surprising the speed at whichtheysweptalong They

rarely ran in a direct course for an}' distance, but dodged aboutamong the bushes in the most bewildering way After runningfor a few moments they would stop and intently regard thecause

of their alarm, and thus give an opportunity to shoot at them

great speed as the parents, keeping ahead with apparently no

such rapidity that they become invisible, and it would seem that

receive on the rough, uneven ground their owner delights in.

to be ratherdependant on a supply of water, though where theindividuals living to the south of Toyo Plain and similar dr}'

of the parietal to the end ofnasals is flat, with aslight slope

either side ofthe median line at anterior end, androunded, being

anterior portion of the maxilla Premaxillae rather long and

slender, nearly straightforthe anteriorhalf, andrising posteriorly

in an upward curve to the nasals Palatines small, produced

Inferiorrimof orbit to alveolus of secondupper molar 50

Madoqua phillipsi (Thomas)

Madoqua phillipsi Phillip's Dik-Dik Native name Sakdro

Phillip's Dik-Dik is larger than Swayne's, and alsover}'

differ-ence in colorbetween thesetwo small antelopes is very apparent

A remarkable peculiarity of these little animalsis theimmense

eye and from its jetblack color and considerable size makes a

and as the skull shows a cavit}^ at this point, no one would

ii6 Field Columbian Museum —Zoology, Vol i.

lar deposit, but of course from the greater size of that species, it

is on a much larger scale.

Phillip's Dik-Dik and the preceding species have short zles and slanting premaxillae, very different in appearance from

until we had reached the southside of the Golis Range It may

exist in Guban; Swayne says it is found all over that section,

M. swaynei, andif it is as plentiful as he states it is very strange

swaynei, asI have stated, is found in both Guban and Ogo, and

the same kind of ground and in their ways and actions they are

them actually together, and I doubtvery much if they associate

with each other If an individual ofone species is shot, it isnot

likely that you will be able to obtain one of the other in the

immediate vicinity, although it ma}' possibly be found a mile or

they were mounted on springs, flying at times over low bushes,

get a better sight of the intruder ontheir grounds Their

jumps of the mule deer {Cariaciis macroiis, Say) than of any

with a peculiar physiognomy, caused by the black swelling

naturally associated with a nocturnal animal, and gives the

the daytime from lack of ability to see clearl}'. The iris is alight hazel, which makes the large eye even more conspicuous

Onlythe males of these species carry horns, about 2.50 inches

in length, nearly straight, sharply pointed, and annulated for

Between and back horns

long crest-like tuft of rufous-colored hair. The face narrows

rapidly in front of the orbits, giving the head a wedge-shaped

of the horns The figure of this species in Sclater and Thomas'

flanks are too light. The skull of this species islonger than that

fossa isverylarge and e.xtends beyond the lacrymal to near theanterior end of the maxilla Premaxillac curved posteriorly but

produced beyond the anterioredge ofsecond upper molar

INCHES

Frontoparietal suture to posteriorendof nasals 1.22

Inferiorrimof orbit to alveolus ofsecond upper molar 55

Madoqua guentheri (Thomas)

Madoqua guentheri Guenther's Dik-Dik Native name

Phillip's Dik-Dik it is found throughout Ogaden In size it is

much largerthan either of the preceding species, andhas a very

differently colored coat, but would without these characteristics

be readilydistinguished bv its long muzzle It is not nearl}' sohandsome a species as the other Dik-Diks, lacking entirely the

and

some-ii8 Field Columbian Museum —-Zoology, Vol i.

rarely more than three are started at a time It may, however,

Dik-Dikdoes not seemto differ from its allies ofthe samegenus

The skull shownonthe plate accompanying this paper will give

a very clear idea of the great differenceinoutline thereis between

premaxilla- with their peculiar shape at once attract attention

This species is about one-fourth larger than Phillip's Dik-Dik

The skull is very high posteriorly at about the center of the

supra-occipital and gradually forward to tip of nasals It is widebetweenthe orbits, narrowing slightly to the nasals These last

are very short, being wider than they are long The premaxillae

their midlength, rising with a sharp curvetothe nasals, affording

large and deep, occupying all the lacrymal and extending onto

the maxilla Z3'goma, short, composed mainly of a process

from the squamosal overlapping a short process from the molar

bullae large.

INCHES

Gazella pelzelni (Kohl).

//. 9 ad. Hullier

This is the gazelle of the lowlands and is not often seen much

habits Pelzeln's gazellefrequentsdryandstonyplaces, covered

it inhabits It goes in small troupes of from two or three to

number I ever saw togetherat one time As a rule, it is not a

wild creature and readily permits an approachsufficiently nearto

straight andannulated nearlytothetips. Thefemalealso carries

indi-viduals and I hardly know what causes it. The typical stylehas

a broad conspicuous chestnut band running lengthwise along thebod}'just above the white of the bell)^ But some individuals,

distinguish-ing mark It may be possibly an exhibition of individual

was this peculiarityconfined toeither sex.

The skull of this species is rather deep through in comparison

with its length The nasals are rounded ontop and bendslightlydownwards at tip. Theyare also very broad and of about equal

width throughout their entirelength The occipitalregion curves

interparietalwelldefined,itslengthbeing nearlyone-thirditswidth;

I20 Field Columhian Museum —Zoology, Vol i.

its width to end of nasals, whereit narrowsrapidly to endof

small, and not produced much beyond the middle of last molar

nasals, which is not the case with the others mentioned

INCHES

Fronto parietal suture to supraoccipital 2.50

Circumferenceofhornatbase 3 60

Gazella spekei (Blyth)

This is the gazelle of the high plateau, and first appeared to

seen in the vicinity of this locality, but Pelzeln's gazelle was not

found any farther to the south, the present species replacing it

be confounded with it, as it is easilydistinguished by its peculiar

the female It inhabits the same kind of ground as Pelzeln'sgazelle isfound in; stony bare stretches with patches of stunted

trees of any size, although I have known it go into rather

dense clumps ofjungle I think, however, on these occasions it

than from choice Bothsexes possess horns,thoseof the females

are rather stout, deeply ringed almost to the tip, and curved

gazelle It is ashy animal, attimesquite wild, and oneisobliged

as if the malesgenerallyled thesmall bands inthemarchesabout

antelope But of course there were instances when I saw the

a very careless sort of way All the individuals, however, keep

a pretty sharp watch, and when anything unusual is seen the

whole band stops andregards it intently, for the animals possess

a good deal of curiosity and frequently stand and gaze a long

commence to graze again, or play with each other, the males

burst of speed; for they run with much swiftness If followed

persistently for a time they will leave the locality entirely, at

posterior end resting on the maxilla, which intervenes and

the recesses being quite deep Lacrymal fossa wide and deep,

at base, compressed laterally, narrowing rapidly at tip, which

122 Field Columbian Museum —Zoology, Vol i.

IN'CHES.

Anterioredgeof external auditory meatusto supraoccipital i.^g

Circumferenceof horn atbase 3.30Lengthofhorn along curve 10.50

AOld.

j.-n. $ ad. Silo Plain

0. $ juv. Silo Plain

i- 9 juv. Silo Plain

in any numbers until the country south of the Golis Range is

herds, sometimes several hundred are seen together, and

able circumstances, thewild animals must have difficulty in

the sea, the natives get their supply bydigging in the dry beds

of the Tugs, or rivers. Of course the animals can not do this,and they must depend upon the chance pools they meet with,

that is, close enough for a fairly near shot, something over a

not difficult to get very near one of these animals if stalked with

knowledge and judgment

It does not seem to make much difference to this gazelle

trees and bushes, or on bare plains, but I think it prefers the

than in any other But it can not be considered as distinctly ananimal ofthe plains, as for instance the Hartebeest is. Aoul can

Possessed of considerable curiosityAoul will always stopafter

large for a gazelle, the body presents but a small mark, and thebullet must be placed in somevital part or else the animal will

seem to incommode it in the least, nor prevent it from running

at full speedon the three that are unhurt

however are much more shapely than those of the extreme

lengths, the latter losing much of the characteristic gracefulcurve

124 Field Columbian Museum —Zoology, Vol ị

and we met with them ahaiost everywhere, being very plentiful

in partsof Ogaden On the Hand they seem to restrict

them-selves mostly to the plainswhere they are seen in large herds

Ammodorcas clarkei (Thomas)

Ammodorcas clarkeị Clarkés gazellẹ NativenameZ>/^d!/ậ

ẹ $ juv. South of Toyo Plain

This rare species is only met with in the country southofToyo

Plain, and then eastward to the landof theDolbahantạ It does

not seem to bevery numerous even in the localities it frequents,

is not easily seen among the bushes, and it has the habit of

top, which its longneck readil}^enables it to dọ The neckisso

other, yet at a distanceit is not always easy to distinguish them

well up, the former inclining slightly towards the latter, clearing

running so that they nearly touched each other, as some writers

have described, but on the contrarythe tail is carried straight up

in the air, and this alone, from its length, would at once

the long neck and peculiar physiognomyof the Gerenuk, both

same

kind of country, butI have never seen them associated together.

As mentioned by Swayne, it frequents thejungle of the umbrellamimosa, and glades of dur?- grass, often taking refuge in dense

of the male are peculiar and shaped more like those of a

beautifully colored animal, and possesses more sporting qualities

species of Koodoo, particularly the lesser Koodoo

Roofof cranial cavit}' convex from rear of pedicles to

Inter-parietal rather large; facial portion flat, descending rapidly atthe

posterior part of theorbits,andretainingitswidthtothe posterior

anteriorly Lacrymal fossadeep and occup3'ing the surface of

the nasals, where the articulation is very broad Pedicles only

moderately high, the base of the horns elevated slightly above

long, curved inward at tip, and hanging well away from andbelow the auditory bullae. These last are onlymoderatelj'^ large,

rather short, broad above, ending below in a hamular process

widen-ing towards the pterygoids Mandible long and slender, in this

respect resembling that of Lithocranius walleri; the condyle isbroad andflat, the coronoid process long and narrow, and curvedbackwards, anglepartiallyroundedextendingconsiderallybeyond

small, molars large, occupying more than twice the space of

upward

126 Field Columbian Museum —Zoology, Vol i.

Circumferenceofhornsatbase 5.25

The skull of this species is similar to that of Waller's gazelle^

prolonged according to its comparative length, and is altogether

of a more delicate shape

Lithocranius vi^alleri (Brooke)

/-•. $ ad. Mandera

also the most peculiarin appearance, the exceedingly long neck,

large eyes, lengthened muzzle and general shape of the face,

when startled drops itshead on a level with the body and sneaksoff among the bushes, in a verydifferent manner fromthe gallantwayin which the Dibatagremoves himself from the object of his

fears.

The Gerenuk goes in small troupes of from three to eight or

ten individuals, and is found usually on rocky ground broken

with ravines and covered more or less densely with bushes and

Avhere grass is partly orwholly absent It has great curiosity,

which failing is often fatal to its safety, and when it sees anunusualobject, a hunterfor instance, will stand andgazesteadilyfor some moments without moving If one is not near enough

toremain absolutely rigid until the Gerenuk commences to feed

the head isducked behind the bush and the animal sneaks away

approach to a fair shooting distance could begained Iconsider

the Gerenukas rather astupidanimal,certainlyitdoes notseem to

careof itself whichare common attributes of allother species ofgazelle The meat is poor, being like the flesh of allothergame

curve The females are hornless The Gerenuk is never seen

on the treeless plains, such as aremet with ontheHand, at least

on barren, rock}^ hillsides and summits, the valleys betweenwhich, however, were covered b}' a thorn forest. In such cases

another The usual gait of this antelopeis aslouching trot,with

when really frightened it gallops with considerable speed, ping, however,at intervalsto look back at the objectof itsalarm,

forgets its fearandcommences tofeed orwalk slowlyalong Themale of this species stands about 4 feet 6 inches high, a consid-

128 Field- Columbian Museum —Zoology, Vol i.

side of the face awhitish stripe between theeye and end of nose,

their brethren in other parts of the countrytothenorthandwest

I was impressedwith this peculiarity, andshouldhaveconsidered

section of the country, but on careful examination of the

found onewith this stripe indicated along the nose, but not so

clearly defined as is generallythe rule in the individuals fromthe

gazelle, as both have similarly shaped faces, very narrow and

pointed Taking the extremes of the two styles, the strongly

marked white stripe, and its almost total absence, one would

up the antorbital vacuity, a black secretion which forms a large

conspicuous prominence on each side oftheface. Theonlyother

removed from the animal, nor the skull, give any indication of

drawing given of it, presents no resemblance, so far as the face

in the museums have this part laid flat to the skull and painted

white, being more of a caricature of the live Gerenuk than

if not in movement,and having an extraordinarystructure of face

and peculiar expression, which cause it to be a rather unique

species among its allies in the family

Theskull of this singular gazelle is remarkable among those

of its relatives for the large proportion that is exhibited behind

the horns, caused by an elongation of the occipital region The

interparietal isvery long, the length being almost equal to the

Between

there is a great depression, causing the facial region to be on a

short and narrow Thenasals are very long and pointed pos

the nasals, but this is not a unique character, as it is seen in

small, but long for their width and slightly compressed laterally.

The mesopterygoid fossa is narrow and very long, while the atines are comparatively short The tooth row is short and the

pal-teeth small and of the brachyodonttype Theorbitis very large

and surrounded by bone, and there is a slight antorbital

on a line with that of the rami, anditsposterior edgeonly slightly

is a singularly long and narrow skull, the greatest width just in

frontof the horns, and from the eyes taperingrapidlytothe nose,

pecu-liar and unusual expression Thehorns arelyrate,curving

15 and 16 inches, but such horns are rare. They vary

inwards toward each other instead of forward

INCHES

Posteriorendof nasals to fronto-parietal suture (median line) 2.40

Lengthof skull fromfronto-parietal suture to supraoccipital 3.45

Centerof external auditory meatusto supraoccipital 2.60 Inferiorrim of orbit to alveolus of third upper molar i.io Inferioredgeoframusto alveolus of firstlower molar 70

130 Field Columbian Museum —Zoology, Vol i.

Oiyx are not often seen in the country north of the Golis

Range, but their numbers increase as the Haud is traversed, and

on the south of Toyo, and in Ogaden they are plentiful It is a

stoutly built animal, about the size of a large donkey, and goes

in herds, sometimes as man}' as fiftyormore being seentogether

is not unusual to find old bulls wandering about by themselves

then become smooth, and end in a sharp point The average

longer, and the cows' horns sometimes j-each a length of 37

or 38 inches They are ver}- formidable weapons and it is

lance-like horns are quite capable of passing entirely throughthe

and are often seen many miles from any place where it could be

Toyo, Silo, etc., in herds sometimes of large size, and subsist

of a bull is very thick on the neck and withers, in some cases

as much as three-quarters of an inch through The natives select