Giáo trình các nguyên lý về bệnh hại cây trồng -chương 8 potx

gi¸o tr×nh gi¶ng d¹y trùc tuyÕn















Gi¸o tr×nh

C¸c nguyªn lý vÒ bÖnh h¹i c©y trång

Ch−¬ng 8

dÞch bÖnh cña c©y trång

Biªn so¹n bëi PGs Ph¹m V¨n Kim

Bé m«n B¶o VÖ Thùc VËt Khoa N«ng NghiÖp

§¹i Häc CÇn Th¬







+
0
/

+"

*
+%$
$
+

 
*
12
3(
"
$%$

/
%+*
4
/
&
#
/
+%-

5"
"

1















1








1
6



"
%7

1
8
""
1+
"(


1
8
""
1+
+


16
8
""
1+
'49
 





6

=



 #$%#%&'#()%*+,-.(/

>
'
""
1+
1



 -
?%

((

/
"
'/

;+
"*

!%

$*

!%


"*
1#* -=
10
$%
%*
"#$
@%
&
@(
$
&+
!%
*

&

(
A#
'
12
2

+
""

1+
+

A
1+%-
""

1+
"(



 

#
!+%


&3

'

(
/

3
/-
5
"

#
+%
)+

+"
(
A#

((

+

!
&/
A)0
#
+%
A/
"
4
(
A#*


(
/



"
4-
B
(
A#

((

A)0
1
1#

 0$%#%&'#('#1#1

5"#$
)+

A
""
1+
"(


'
""
1+

+
-
>
'

'%
(

1+
-
C"
 

10
"#
'
1#
+
1"2(
3
49
1
"0(
@#
&
2
A
""
1+
/%
-

 /$%#%&'#(2345-6.

:
!"

'
1#
+

1"D(
 


2
A
""

1+

'49
 -
E
""
1+

 

/
'"*
!
"#$

'

/0
&
"0(
(/%

"
"

+

-



2
A
""
1+

'49
 
+
%



A
17

<
+%
2
+

$
'%
+
(
10



$%*
((

!
+
10
/%

4


$"
1"
&+

'49
 -


((

/0-

<

0
+%
2
+


"
10
$
#
1

'+
12

+4
4
/

*
((


"
"
%
F%
"*
(
A#
"0(

#$*

 


!

G

   %*+.;%*(

  

 =%>?()

  &@A%;#B5







  

"%*+.;%*(5'#(%5%CB%

%
#
#
+"
"%"
+%

/
'

A/
3

&
A/
$%
"2

 


'7

 #$%*()&@D/.E#;@5;F'3(//.E#&?A)

!"
"

(%
;4 4

4*
+
/
#(

J

4

(D"
+

@#
1#
4

/%
'
+%
!
4

+

1#
K
&#


&3

&#-
/
&

+
1"
!"
'

*
%

&

+%
#(
&
D(
/%-

5+
!"
1+%*

&
'*
&"
!2

,

"
-

 0$G,&@D/.E#;@5;F>#1?H,&@D/.E#&?A)

!"

"#
12
+
((

A"
A0

&
@(
"D(

&+
-


#
%

%
+%
A/
(
&
1(*
%
*

&

'
4N
4(-


+
(
(/*
D(
/%-
5+
!"
1+%
#

"
49$
+
(
!




/
!
@
-

%
'+"


*
/%
'


/0
&
1#

A)%
4
!+%

+%
/%-
>

'

 -

5,
7

M
#
0


"
&+
#"
(

(/

%
H
>M:?

+

#(

+
#"
(

@


*
"
(/
(

@
A#
"

'2*
'"
(
(/
/%
A
/

?+

+

/%
+
4(-

>#

&

/
+%


$

'
A)%
4*
!+%

+%
/%-



/
#
,

+%
"
$


K

4N
'+-
+%
/0
&/

"
"
1#
PIL
-

#

$%
&'
&(

)

&!%
*
+

)
"%
,
"

'-%
-
%).
)
$

,/
+
0)
%1
%

,'
%

2 34
-

5
%

!


4

9
4
-

%

!'
5

&
;

$)

0

+<


*
+)

.
%!
=
+)



)
)
%

%
&

"
+-
)

+-

)
6


)
)
>
%4
)

)

$%?

$-
9B4888
+
C
%
$%
&)
)


&6


&-
+1


)
5
1
<


!'

$%-
#

+#
*#
)

"(


&)'
%
"
)
)
"-.
D-.
&-


$-
9B4888
+

C
%
=
)
)
&%1

6


,'
%
.)
(
-
)
"(


@
A
)
>?
E

%
-
+
)
)

&6

&-

+/

@
A

)
B?
E


%
-
+
5

=
)
)
5
&%1


2%

)
*-
F%

$%?

−



E
)
(
-4

C#
F

,
"(

/%
("

3



+

+
/%
'
"
'
1"
!"

"#
+

A/

$%

A"

&

$%
"2

 

)%+
0

"*

Q

*

/
!"
1#
!

+
+
/%
'
"*
#(


0

!

#

$%
A"
&
$%

"2

 
-
+%
2
+%"*
+
"
"
'
'
+%
(4

'%
+
 

0
@
-


H
@


!"
1"
!"
+"

!

R
'%
%

@
!2
+
1#
,
$(

&"
$


+-
5+

!"
1+%*
$
'%

#(
&
&"
!2



+
1#
'"
,
("
/

-
R
'%
%
#

+
1#
,
$
+
!+
J-I
1#
K*J-

"2

#(*
&"
!2

&

#

+
1#-
B"
+%
&"
12
1/
$


1#
'

&%"

+
A)0
+%$
$

#
%
 

+
1#-

M
&

'%
'%

/
$%

"2
#
4

/0
!

&/




)

!#*

5"
"

"#
A%
+


"


49
0
+%


/

1#


-
5,

7
1"

!"


"
&
"
+
/
/%
#
A/

$%
"2



)%+
0



+
#(
(
*"
S-A$-
!"
&


1#(
'%





+


49

&

"
/

(
)-

"
+%
A/

(
-
%
#
#

%

12
A/
$%
"2

-

5
)+
"

5,
7
CT

&
UA
;PQP=
"
/%


'%
(

#
#

'%

P

"
+%
0
F#

1 
A/
$%
"2

2
&
A/
#$


/
'"


*
0

'
%
+%

/
'
(#"
/
%
"/0

&
((
-




+
1#
QL
-
/
&*

!"
!,
V

,
/%
;"#
+%
/%=-



1

"

/%
#
A/

(
A#

%-
5/
3
%
!
#
(




+


+%
2

#
A
G
"
+

1"
!"


1
V
+-

6I


F

((

+

+%
#
"

+%(
A"
&

!%

A#

/0
#
!

/
3

'
((
*
/

0
+%
2


"
+
F%

3
$%
"2


((
*
&

0
+%
2
%

12
A/
$%
"2

((

-
+
1+%
("

/
A"
&


F

*

!
0

/

+
1#*
'
#
#

+%$
$

/
'
A/
$%
A"
&
$%
"2

 
-



"#
"
/



A
#*
#
2

"
'+"

#

-
B%"

%$
'/


((
*
#



A
"
/*

A)0

"

4

A+
&%"


!+)
(
+

1
10
"

/
-
5,

7
M
(
!"
'%


"#
$
-
0

/
/
/
&*
)%+

0


+
#(

5+
!"
1+%*
&"
+%
$
+


!

1#"
0
"%$
+


+%
2
$%
"2

 -

5+
%
'+"
$
+%*
$

+%
&"
+
#

F

+%
%
#
+


&
'(
(
'"
F%
3
,

%"-
%
((

#



%


A)0
D

@(
$
&+


+
&%

#
+




/

$%

"2


&
A"
A
0


-
.+
1+%
(
A#
((
1"
!"
!%-
5,
7

M
%
'%
'%
+

#(
Z[R"'"
+T)Z"
A)0

"


"
"

4
-

6

?


?
((


C
2
@


+%

/
!

 

C
2
@




A)0
@
-

"

+%(
A"
&
1#"
!%-

 P?N)M@+5>#1,?+,0*(



'
'
+%
A/

#
1#"
&%"
((
-
5
"
A/
#
1#"

S+
<
))
+)$=

\'+-


!
&/

1+%

#
'
"#
"D(
&%"

((

1
'/

3

 


@


A#
,
"
,

&%"
"#
"D(
(
"-

+

A"
'%


((

&%"
"#




A)0
#

4
)0-
"#
!%


H
$%

,
!%

(3
&%"
+

A"
'%
/
/%

((
-


3
3

#
A)0
2

12"-
.+
1+%

&"
A/

"#
!%

0
H
'

/

1/
&%"

#
#

+"
"%"

+
&"

3

 
-

6


"
3
A)0
!
+%
"#
!%
-
?%

A/

#
1#"






&

((




/%

#

A+
&%"
#

#
+"
"%"
+
&"
3


 
-

 Q=%>?()&@A%;#B5

$%
"2

((

&
'(
+
 

!
2
@
*
/%
'
%


'
+%(
A"
&
1#"
!%
-
.+
1+%
#
%
#
#
!%

("
/
!

 

$%
A"
&
$%

"2-
5,
7
B+
4(
PQ*
"

]))'



(/
+
&
!)%+
"
'(
+
1#

+

3


+
+

/%
'
"*

+
1+%

 

#
%



4N
%
+%
"
$


(
+
&/

>#"
&%"

 

)%+
0


"
"
>M:?

3
"
$%$
+%
"

$


10



+"
(*
0

'
"
$%$
/
"-
M"
&3
!"
+%
"
$

@
!2
&
%
&"
A"
&
1#"
!%
!%
#
1%
!2*

10
A/%

#/
A/
$%
"2

66

 "S'155I1#-6/0*(

<

:/



"#

#*
!

+%
/%

((

12
")+
-
#

!

<
M+%
$
/

+



&/
"%$

$%
"2
#
&/

#$
&

"
12
/
 

+
1#
;A+"'<+)
"A)A)A=-

<
%
"
$%$

%
/7
'
*
@)
*


'(

+
4

<
5+%


#
&
/%"
/%
1
10

!"

"#
0
'
"
$%$

"#

6G

<
:/

"#


+%
(
")
!%

/
/%-

5,
7



'
@+49

'%

'%

+

&"A

aa:B

;")
)
A
&"A=



10



 

"
H

5"
"
&+
4(

PQP-

>2

$+
/




+

%
4(
A*
%

10
%$

"
$%$


#

"#
'%

"D(


4N
"#
'%
!%

;[a
6K=-
B%"
"
$%$

%

10
"0(

#$
%$
'/



@#
;

'

("

"%"



'
@+49
'%=-


H

5*

:+%
54*
BH
?+*
5
B"
&
>
5%$*

"
"
"

4(

#
'%
+
&
1
@
PP<PP6
;H
"
H

5

"
"
'
1#

II-III
#
'%=-
M"

$%$
$+
/
 



+
&
)

PP6

,
/
/%
2


"2
"#
&


12
$+
#


#*

!
12
(
A#


'
+*









 P@A%J'B>CB%-6/0*(;%&#U0@D/J#B)

"#
%
1#"


$+%
+


"#-
/%
2
%
A/

#
)+
#
1%
12

1/%
A/
$%
"2


 
-


A)0
$
#

!"


6J

#

14

12

1#"
$+%-

"
#
+
%
&

+
)
+4
(%"
@#

3
$"
"

A%(
12
"
"

-



 "#B/2'#(%-I(0#B'-6/0*(/.E#/?N)M@+5

+%

'+"
/
%+
 
7

#$I(0#B'-#1%#(7

'

/

%+

+

4(

4(

+4
(/"
4(
%"
+

 
A

+
/%

"
4(
F
&
3
3
$%
"2

A


"#

&

"$
&


&

"$
+%
'"
F
1#

"$-
5,


/
!#
+
'$
%


(%
A
@#
#

"$*

(%
A
@#



+
&
/
&*
&&
-

0$I((0#B')M.5#(
'

/
%+

3

3

 

A

+%
2
@



M+
B
5/
B
%
H
/
"
12
'$
!#
+
A
@#
+
&
%"*
+
1+%
/
%+

0
"%$



'$
!#
+

!#

A%
&%"




"#

A

,
1%
+
/

(
&
+4
+

"
(
&
'"
+
&%"
-

./
%+
'+"


H
+
0
+%
A"

'(-
5,

/
&+
&
1
@

&%"



0
/
%
A/


#



)+
#
1%-
?%
#
"

C+

@0
*
)+




:


"$
H
U
"*
12
/%
2



%
1#"
$+%
&%"


*
"%$
$
49
 
! $
"-



"
%7

1
8
""
1+
"(


1
8
""
1+
+


16
8
""
1+
''49
 



...


)
)
>
%4
)

)

$%?

$-
9B 488 8
+
C
%
$%
&)
)


&6


&-
+1


)
5
1
<
...


!''

$%-
#

+#
*#
)

"(


&)''
%
"
)
)
"-.
D-.
&-


$-
9B 488 8
+

C
%
=
)
)
&%1

6


,''
%
.)
(
-
)
"(