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The Encyclopedia Of Nutrition And Good Health - I pps

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The stress response is modified by hormones, and sustained, elevated adrenal hormones due to prolonged stress affect the immune system and decrease the immune response to foreign invader

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356

iatrogenic malnutrition See HOSPITAL-INDUCED

MALNUTRITION

IBD See INFLAMMATORY BOWEL DISEASE

IBS See IRRITABLE BOWEL SYNDROME

iceberg lettuce See LETTUCE

ice cream A frozen DESSERTcontaining flavored,

sweetened frozen cream and MILK products, and

SUGAR Frozen desserts based on EGGS, cream, and

milk were apparently invented in the 1600s,

although frozen blends of fruits were served in

ancient China, where salt mixtures were used to

lower the temperature below freezing Americans

eat an average of 15 quarts a year per person

Ice cream is a high-CALORIE, high-FAT, and

high-sugar food Unless it is homemade, ice cream will

usually contain artificial coloring, flavorings, and

stabilizers such as LOCUST BEAN GUM, GUAR GUM,

CARRAGEENAN, GELATIN, and alginic acid or cellulose

derivatives Federal law requires that ice cream

contain at least 10 percent fat (BUTTERFAT), and

most regular ice creams provide 10 percent to 12

percent butterfat Super-premium vanilla ice cream

contains 16 percent to 20 percent butterfat In

con-trast, ice milk contains 2 percent to 7 percent fat,

while sherbet contains 1 percent to 2 percent fat

Ice cream usually contains sugar or other

sweeten-ers and emulsifisweeten-ers like POLYSORBATES and MONO

-GLYCERIDESstabilize ice cream during processing

Nondairy frozen desserts resembling ice cream

have no legally defined butterfat content, and

although they are low-fat options, sometimes these

desserts have more total calories per serving than

traditional ice cream due to their high sweetener

content

One cup of rich vanilla-flavored ice cream with about 16 percent fat provides: 349 calories; protein, 4.1 g; carbohydrate, 32 g; fat 23.7 g; cholesterol, 88 mg; calcium 151 mg; vitamin A, 219 retinol equiv-alents; thiamin, 0.04 mg; riboflavin, 0.28 mg; niacin, 0.12 mg

One cup of regular vanilla ice cream with about

11 percent fat provides: 269 calories; protein, 4.8 g; carbohydrate, 31.7 g; fat, 14.3 g; cholesterol, 59 mg; calcium, 176 mg; vitamin A, 133 retinol equiv-alents; and similar amounts of thiamin, riboflavin, and niacin to those found in rich vanilla ice cream One cup of vanilla-flavored ice milk with about

4 percent fat provides: 184 calories; protein, 5.2 g; carbohydrate, 29 g; fat, 5.6 g; cholesterol, 18 mg; calcium, 176 mg; vitamin A, 52 retinol equivalents; and similar amounts of other vitamins (See also

DAIRY-FREE FROZEN DESSERTS; DIETARY GUIDELINES FOR AMERICANS.)

ideal body weight An outmoded standard for body weight that originated from data collected by U.S life insurance companies from their policy holders who lived the longest Ideal body weight was listed as the average weight for a given age, height, body, or “frame” size From these data grew the classic definition for OBESITYas a weight

20 percent greater than the ideal body weight Part

of the problem with this standard of ideal body weight lies in the nature of HEIGHT-WEIGHT TABLES Weight charts and tables do not take into account individual variations in body structure (muscles and bone mass); consequently, there is no stan-dard for measuring small, medium, or large body frames

There is no single ideal body weight for a group

of people because every person has an individual, desirable weight Selecting a realistic body weight

as a goal has proven to be far more useful than

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striving for a rigidly defined standard A realistic

body weight is one that can be readily maintained

without intermittent DIETING

A measure of body FATis more useful than total

body weight because body weight may not

corre-late with the amount of fat on a lean person

Fur-thermore, ideal body weight provides no indication

of fat distribution Some fat deposits are riskier for

heart disease than others Methods of accurately

measuring body fat at clinics include buoyancy

testing with underwater weighing (requiring a

spe-cial tank) and SKIN FOLD thickness measured by

skin calipers or by skin resistance measurements

Suggestions for estimating an optimal body

weight include: For males, take 106 pounds for the

first 5 ft of height, then add six pounds for each

additional inch of height For females, take 100

pounds for the first 5 ft., then add five pounds for

each additional inch (See also BODY MASS INDEX;

FAT-FOLD TEST.)

idiopathic A medical term that is applied to a

disease or condition arising spontaneously from an

unknown cause Idiopathic GOUT and high blood

pressure are examples

ileitis See CROHN’S DISEASE

ileum The last three-fifths of the small INTESTINE

that joins the large intestine (COLON) The length of

the ileum varies among individuals, ranging from

15 to 30 ft in adult men The ileum absorbs FAT,

fat-soluble vitamins, CALCIUM, MAGNESIUM, VITAMIN

B12, and AMINO ACIDS BILE salts, which act as the

detergents in bile required for fat DIGESTION, are

also absorbed in the ileum and are recycled by the

liver to released once again in bile

Flow of material between the ileum and colon is

regulated by the ileocecal valve (See also ENTERO

-HEPATIC CIRCULATION.)

illness Ill health or disease; the opposite of

well-ness Illness reflects imbalanced body functions and

can thus be regarded as a change away from the

healthy state, in which all systems function within

normal limits (HOMEOSTASIS)

An illness may be localized, in which a limited

region of the body is affected, or it may be systemic,

in which several parts of the body or the whole body are affected Pain is often associated with illness, although pain is not equivalent to disease Pain gen-erally indicates that an imbalance exists in the body The imbalance could reflect an unhealthy lifestyle, such as dietary excess or deficiency, or it could be associated with infection and inflammation Most illnesses are self-limiting, meaning that the body generally can cure itself when given the oppor-tunity It is now clear that the body’s systems work together to maintain health Particularly important are the IMMUNE SYSTEM, the NERVOUS SYSTEM, and the

ENDOCRINE SYSTEM (hormone-producing system)

An imbalanced immune system affects the brain, and the brain alters immunity and hormone pro-duction Hormones in turn affect nerve function

As an example, perceived STRESScan trigger a FIGHT

OR FLIGHT RESPONSE by the brain The stress response is modified by hormones, and sustained, elevated adrenal hormones due to prolonged stress affect the immune system and decrease the immune response to foreign invaders

Whether or not an individual becomes ill depends on a complex interplay of many factors, broadly categorized according to medical history and environmental influences Family history reflects patterns of inheritance, over which one has

no control Genetic predisposition thus increases the risk of many chronic, degenerative diseases of

AGING, such as OSTEOPOROSIS, heart disease and

STROKE, high blood pressure, and DIABETES Health history can have a profound effect on susceptibility

to illness Thus, prior injury, deficiency, or illness can set the stage for a subsequent illness For example, arthritis can begin at the site of a former injury, and prolonged treatment with a broad-spec-trum antibiotic can destroy beneficial gut bacteria and promote a yeast infection Prior drug treatment can alter the body’s ability to destroy alcohol, and pretreatment with alcohol can alter the body’s abil-ity to degrade many drugs In terms of public health, immunization against polio, flu, or tetanus reduces the risk of these diseases At the beginning

of the 20th century, niacin deficiency caused wide-spread pellagra and associated mental illness in the South The enrichment of bread and grain products with the B vitamin NIACINin the 1920s essentially eliminated pellagra as a public health issue in the United States

illness 357

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Due to differences in inheritance and

environ-ment, toxic exposure, nutritional status, and

med-ical history, each person is biochemmed-ically unique

Thus, individuals vary in their ability to protect

themselves against damaging effects of FREE RADI

-CALS, and to detoxify potentially dangerous

chemi-cals that they eat or drink or breathe at home or in

the workplace

Although neither a family history nor a medical

history can be altered, lifestyle choices can

pro-foundly impact susceptibility to illness The most

frequently cited RISKfactors for illness and

prema-ture death include the use of tobacco and alcohol,

accidental injury, unwanted pregnancy, drug

abuse, and inadequate nutrition The growing

awareness that prevention is the most

cost-effec-tive and permanent solution to many health issues

has lead many physicians to work with their

patients as partners in health to empower them in

making healthful choices for themselves and their

families

This approach could possibly decrease American

deaths before the age of 65 by two-thirds, even

without further breakthroughs in medicine and

nutrition

Prevention and Personal Responsibility

The health-conscious individual can focus on four

essential steps in maintaining health and preventing

illness: positive attitude, a healthy diet, regular

exer-cise and minimizing toxic exposures Humanistic

psychology emphasizes the importance of a positive

mental attitude in preventing illness and

maintain-ing health The immune system and the repair

mechanisms of tissues are well designed to ward off

infection, to cure disease and to repair injury

DEPRESSION seems to diminish this capacity For

example, depression causes a drop in the production

of interleukin, proteins that help regulate the

immune response and help activate cancer-killing

lymphocytes Clinical statistics in the United States

suggest psychological stress can harm the heart,

increasing the risk of rehospitalization among

patients with cardiac problems Group therapy,

meditation, and other practices may improve

car-diovascular health in patients with clogged arteries

by modifying their response to psychological stress

Susceptibility to disease reflects nutritional

sta-tus; the nutritional environment affects the

expres-sion of inherited traits Either overnutrition or

UNDERNUTRITION can set the stage for chronic ill-ness Excessive fat consumption increases the risk

of cancer and obesity, while inadequate amounts of most nutrients eventually lower the body’s defense against disease, including cancer On the other hand, a healthy diet provides optimal amounts of all nutrients to assure health, while avoiding detri-mental food constituents

Probably the best strategies to avoid chronic ill-ness associated with aging are regular physical exercise coupled with wise food choices A seden-tary lifestyle is linked to an estimated 250,000 deaths annually in the United States Exercise decreases the risk of heart disease, stroke, high blood pressure (hypertension), adult onset dia-betes, osteoporosis (thin bone disease), and colon and breast cancer

Minimizing exposure to potentially damaging agents such as solvents, pollutants, and cigarette smoke is also important The amount, type, and length of exposure to a toxic material at home or in the workplace affects health Only recently have studies been undertaken to determine additive, long-term effects on health of continued low level-exposure to pesticides and industrial pollutants in food and water and air (See also ANTIOXIDANT; BIO

-CHEMICAL INDIVIDUALITY; DEGENERATIVE DISEASES;

DETOXIFICATION.)

imitation fat See FAT SUBSTITUTE; OLESTRA; SIMP

-LESSE

imitation flour See FLOUR SUBSTITUTE

imitation food A processed food that is nutri-tionally inferior to the real food The designation

“nutritionally inferior” on a food label means that the food contains 10 percent less of one or more nutrients than the food for which it substitutes The food industry has devoted considerable finan-cial resources toward producing imitation food, most often substitutes for MEAT, FISH, dairy prod-ucts, and FRUIT For example, SURIMI is imitation seafood Compared to real foods, imitation foods often have a longer shelf life and may be tastier Food additives are carefully selected for this pur-pose Imitation foods are often less expensive than

358 imitation fat

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the real foods, but are generally less nutritious than

the foods they replace because processing destroys

or removes important nutrients Relatively few

nutrients are added back

Current regulations specify that food labels must

list ingredients in descending order according to

weight, meaning that the first ingredient listed is the

most predominant If the first item listed on the food

label is one of the following, the food is likely to be

fabricated or highly processed: any natural

sweet-ener such as corn sweetsweet-ener, high FRUCTOSE CORN

SYRUP, HONEY, DEXTROSE, SUCROSE, or corn syrup

solids; TEXTURIZED VEGETABLE PROTEIN; and sodium

caseinate The presence of artificial food colors, MSG

(MONOSODIUM GLUTAMATE), COCONUT OIL, palm or

palm kernel oil, and PRESERVATIVESalso indicates that

the food is imitation or highly processed (See also

CONVENIENCE FOOD; FOOD ADDITIVES; FOOD LABELING.)

immune system An elaborate, finely tuned

defense system to destroy and counter the effects of

viruses, bacteria, yeasts, and foreign substances

that operate within tissues and cells and in the

bloodstream The immune system recognizes “self”

from “nonself,” substances not part of the body

Another feature of the immune system is memory

It can remember previous invaders and mount a

rapid response to them when they reappear When

the immune system is healthy, it destroys foreign

elements without causing symptoms, but an

imbal-anced immune system can set the stage for disease

Foreign substances and microorganisms may not

be recognized or destroyed, resulting in chronic

infection or even CANCERand AIDS An imbalanced

immune system can attack the body’s own tissues,

creating AUTOIMMUNE DISEASES, or it can

over-respond to common substances, creating allergies,

such as food allergies

Organization of the Immune System

The two branches of the immune system are

“cel-lular immunity” and “humoral immunity.” The

first depends on the active participation of different

types of cells Cellular immunity includes

macro-phages, cells that engulf foreign invaders These

scavengers are amoebalike cells that surround and

digest foreign particles, viruses, and bacteria

Macrophages live in tissues like the spleen

(spleno-cytes), the LIVER(Kupffer cells), the lymph

(wan-dering macrophages), the spinal cord, the brain (microglia), and connective tissue

Lymphocytes are an important type of white blood cell T cells are highly specialized lympho-cytes that attack viruses, tumors, and transplanted cells and regulate the immune system T cells are processed by the THYMUS GLAND They work with B cells, which produce defensive proteins called ANTI

-BODIES In a typical scenario, macrophages first engulf foreign materials called ANTIGENSand trans-form fragments of the antigens for display on their cell surfaces Certain T “helper” cells, acting as

“generals,” “read” these antigens, and in turn stim-ulate the production of specialized T “killer” cells, foot soldiers that destroy abnormal cells or foreign materials The gut is the largest immune organ, which is called the Gut-associated lymphoid tissue (GALT) GALT produces more antibodies than any other tissue in the body

The different types of immune cells communi-cate with each other via protein messengers called

LYMPHOKINES For example, macrophages produce a lymphokine called interleukin-1 to activate T helper cells, which in turn produce interleukin-2,

to stimulate the production of the killer T cells Helper T cells also produce gamma INTERFERON, which activates killer T cells

Mast cells are a type of T cell that lives in tissues and fights local infection When they contact for-eign materials and cells, mast cells destroy them Mast cells also release special chemicals like HISTA

-MINE, as well as certain lymphokines that trigger inflammation marked by swelling (EDEMA), red-ness, itching, sneezing, and runny nose Lym-phokines also trigger phagocytes (macrophages) to destroy foreigners and dispose of signal proteins once they have done their work

Humoral immunity pertains to blood and lymph It relies on cells that release defensive PRO

-TEINS called complement and antibodies into the bloodstream to fight infection Antibodies (gamma globulins) are Y-shaped proteins designed to target

a particular antigen, by which a substance is recog-nized as being foreign An antibody can neutralize the enemy either by binding to it or by targeting it for attack by other cells and chemicals After anti-bodies bind foreign cells, complement ruptures them Complement also triggers localized inflam-matory reactions, leading to common symptoms of

immune system 359

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pain, redness, and swelling, as well as to an

increased concentration of defensive cells at the

point of injury or infection

B cells, which originate in bone marrow, are a

type of lymphocyte that yield plasma cells,

special-ized to produce antibodies when exposed to foreign

invaders B cell proliferation, maturation, and

anti-body production are stimulated by T helper infected

cells Another type of T cell, called T suppressor

cells, gear down the immune system by turning off

B cell production Thus T suppressor cells limit

aller-gic attacks and auto-immune reactions

Immunity

Immunity is a hallmark of the immune system

The recovery from an infection renders the

indi-vidual immune to subsequent attack by the

partic-ular disease-causing agent Immunity to chicken

pox is a common example The underlying

mecha-nism relies on memory T and memory B cells in

the bloodstream, which signal a red alert for a

quick attack the next time a conquered virus

in-vades again These memory cells multiply rapidly

when they again encounter any antigen they

remember

Nutrients that May Benefit the Immune System

The immune system requires a rich array of

nutri-ents, including protein, FATTY ACIDS, VITAMINS, and

MINERALS, for normal function A JUNK FOOD diet

can lead to overnutrition (too many calories and

fat) and malnutrition (too little trace minerals and

vitamins) that weaken the immune system

Mal-nourished individuals lacking adequate protein or

calories are prone to disease Low-level deficiencies

of many nutrients seem to lower the effectiveness

of the immune system Supplements can boost

immunity, especially in elderly people It seems

clear that a wise approach to support the immune

system nutritionally includes a varied

diet—reduc-ing fat to less than 30 percent of daily calories and

emphasizing whole foods, FRUITS, and VEGETABLES,

especially those rich in VITAMIN C, BETA-CAROTENE,

and other carotenoids Supplementing the diet

with 100 percent of the RDA, (RECOMMENDED

DIETARY ALLOWANCE) of B vitamins and trace

min-erals for insurance may also be prudent when a

diet is compromised by junk food or when the diet

provides fewer than 1,600 calories

ANTIOXIDANT nutrients such as vitamin C, VITA

-MIN E, SELENIUM, COPPER, MANGANESE, and beta-carotene may enhance immune responses by lowering the burden of FREE RADICALS, thus pro-tecting immune cells against the cumulative oxida-tion and free radical attack due to the release of powerful oxidizing agents as superoxide, hydrogen peroxide, and hydroxyl radicals

Vitamin C deficiency lowers the immune response in animal models Adequate vitamin C increases T and B cell production and helps attack-ing cells migrate to sites of infection while makattack-ing viruses and bacteria more sensitive to destruction Furthermore, vitamin C acts as an antioxidant, and

it protects cells against reactive chemicals produced

by mast cells used to destroy foreigners

Vitamin E enhances both humoral and cell-mediated immunity, while vitamin E deficiency contributes to reduce T cells, killer cells, and macrophage function Vitamin E supplementation boosts the immune system in elderly men and women consuming a typical diet, suggesting that older people require more vitamin E than specified

by the adult Recommended Dietary Allowance to assure a fully functional immune system

Selenium is a cofactor for an important antioxi-dant enzyme, glutathione peroxidase, which neu-tralizes lipid peroxide that could damage the immune cells Selenium works with vitamin E to stimulate the immune response to infection in experimental animals Together they may help pro-tect against cancer Selenium increases T helper cells and increases antibody production in experi-mental animals However, excessive selenium depresses the immune system

Other specific nutrients support the function of the immune system:

FOLIC ACID is required for immunity and lym-phocyte production Folic acid is often deficient in the American diet

IRON is required to produce T and B cells Iron deficiency is associated with increased incidence of common infection among children

MAGNESIUM is needed by the complement sys-tem to activate phagocytes It is also required for antibody production Americans typically do not consume enough magnesium

PANTOTHENIC ACIDand VITAMIN B6help keep the lymphatic system and thymus gland healthy

360 immune system

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VITAMIN Aand beta-carotene help maintain

thy-mus gland function during stress Overproduction

of cortisol, a stress-induced hormone from the

adrenal glands, tends to shrink the thymus gland,

which is critical for fully functioning T cells

Beta-carotene is the precursor of vitamin A Limited

studies suggest that beta-carotene may also

stimu-late helper T cells Studies of children in developing

nations indicate that there is a direct relationship

between vitamin A deficiency and decreased

resis-tance to infection On the other hand, excessive

vitamin A can decrease immune system function

Vitamin B6plays an important role in

maintain-ing optimal immunity, includmaintain-ing antibody

produc-tion and phagocytic activity Vitamin B6deficiency

impairs the immune system in a number of ways

It lowers cell-directed immune responses,

de-scribed earlier, and leads to decreased thymus

func-tion Vitamin B6 deficiency during gestation in

experimental animals impairs immune functioning

even in first and second generation offspring

ZINChelps maintain lymph glands and the

thy-mus gland, thereby helping to fight chronic

infec-tion Zinc is required for many important enzymes,

and it is not surprising that zinc deficiency

decreases T cell and B cell function and

macro-phage activity However, too much zinc can depress

immunity Zinc, in combination with other trace

minerals including copper, iron, and manganese,

appears to improve B and T cell function in older

people

Copper deficiency is associated with an

increased risk of infection Copper deficiency

diminishes the effectiveness of the humoral system

in lab animals Copper is an essential component of

SUPEROXIDE DISMUTASE, an antioxidant system, and

CYTOCHROMEC oxidase, an enzyme system required

for energy production

Immunity, Stress, and Exercise

Physical or emotional stress can alter hormonal

output and immune response A high level of stress

increases the risk of illness and injury in the

fol-lowing year and shortens the life span

Emotional well-being is supported by proper

diet and regular physical EXERCISE Moderately

intense exercise increases the production of ENDOR

-PHINS, the brain’s own opiates, which can bolster

parts of the immune system Studies indicate that

interleukin-1 and interferon, which help the body respond to infection or injury, increase after mod-erate exercise Modmod-erate exercise also increases killer cell activity

Strenuous aerobic exercise may decrease effi-ciency of the immune system and temporarily increase susceptibility to illness by increasing the production of adrenal stress hormones Among the hormones produced is cortisol, which limits inflammation by blocking the immune system With continued stress, production of protective antibodies (such as secreted IgA, the antibody that protects the intestine and other body cavities against invasion by foreign substances) Chronic stress also decreases killer cell activity, increasing a person’s susceptibility to disease (See also AGING; B COMPLEX; ENDOCRINE SYSTEM; MALNUTRITION; NER

-VOUS SYSTEM; PSYCHONEUROIMMUNOLOGY.) Chandra, R K “Nutrition and the Immune System from

Birth to Old Age,” European Journal of Clinical Nutrition

56, supp 3 (August 2002): 573–576.

inborn errors of metabolism Abnormal gene products can cause metabolic imbalances, resulting

in disease Generally the inherited defect leads to the inadequate formation of an enzyme Occasion-ally, inadequate formation of a COENZYME(enzyme helper) limits enzyme function or the protein-based signalling system that regulates a given enzyme’s function Most genetic defects are classi-fied as autosomal recessive, meaning that they are not sex-linked Full expression of the imbalance can occur only when both chromosomes contain the defective gene coding for a given enzyme and normal genes coding for that enzyme are absent Examples of genetic diseases related to nutrients include familial HYPERCHOLESTEROLEMIA(high blood

CHOLESTEROL), sickle cell anemia, GALACTOSEMIA, and PHENYLKETONURIA PKU is an inherited inability

to metabolize the essential AMINO ACID, PHENYLALA

-NINE PKU responds to nutritional intervention During infancy and childhood PKU patients receive carefully balanced diets that provide only enough phenylalanine to support growth Labels for foods containing the artificial sweetener ASPARTAME

must warn phenylketonurics because of its pheny-lalanine content (See also DNA; FOOD LABELING;

MUTATION.)

inborn errors of metabolism 361

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Brusilow, S W., and N E Maestri “Urea Cycle Disorders:

Diagnosis, Pathophysiology and Therapy,” Advances in

Pediatrics 43 (1996): 127–170.

indigestion See GASTROINTESTINAL DISORDERS

individuality See BIOCHEMICAL INDIVIDUALITY

induction The increased production of an

ENZYME in response to external stimuli Enzymes

function as biological catalysts of cellular chemical

reactions The body can adapt to a limited extent to

changes in the diet and to environmental

influ-ences by altering the levels of enzymes in a given

tissue For example, a high-carbohydrate diet leads

to increased production of AMYLASE, the

starch-digesting enzyme of the pancreas Starvation

decreases levels of digestive enzymes and enzymes

responsible for fat and glycogen synthesis but

induces enzymes required for fat and carbohydrate

degradation Enzymes that synthesize BLOOD SUGAR

(glucose) from amino acids are also induced by

starvation

Excessive alcohol consumption induces liver

alcohol oxidizing systems so that alcohol will be

cleared from the blood more efficiently Similarly,

many medications induce liver enzymes

responsi-ble for drug degradation Certain cellular enzyme

levels increase in response to hormones and to

growth promoters Examples include the

hor-mones ESTROGEN (female sex hormone), CORTISOL

(adrenal hormone regulating energy metabolism

and degradation), and GROWTH HORMONE (a

pitu-itary hormone involved both in growth and

main-tenance of tissues, especially muscles) INSULIN

provides the broad impetus for increased enzyme

production in many tissues Insulin from the PAN

-CREAS is perhaps the most general anabolic

hor-mone; that is, it promotes enzymes leading to the

accumulation of protein, fat and carbohydrates

(See also DETOXIFICATION.)

infant formula A manufactured food designed to

nurture the infant during the first year of life, until

weaning Commercial infant formulas are either

nonfat cow’s MILK-based or soybean-based

Com-mon formulas are available in powdered form, as

concentrates, or as ready-to-feed liquid with no

prior preparation No formula exactly reproduces human milk; on the other hand, formulas can pro-vide adequate nutrition for babies

In the late 1970s, production of chloride-deficient formulas caused delayed speech, slowed growth, and poor muscle control in babies who had consumed the products Partially in response to this disaster, Congress passed the Infant Formula Act of 1980, which mandates the U.S FDAto see that this synthetic food meets nutrient standards based upon the American Academy of Pediatrics’ recommendations to assure infant growth and development

In 1982, the FDA adapted quality-control proce-dures to monitor the production of this food As a result, infant formulas are nutritionally similar though not identical to BREAST MILK in total pro-tein, total fat, calcium-to-phosphorus ratio, energy content (calories/100 ml), content of the essential fatty acid linoleic acid, and electrolytes (sodium, potassium, chloride)

There are advantages to infant formulas There

is no limit to the supply, and a mother has greater freedom to care for other children or to return to work Other family members can participate in feeding sessions, developing the warmth of that association The mother of a formula-fed infant can offer the same closeness and stimulation as the

BREAST-FEEDINGmother

In 1998 an estimated 29 percent of women in the United States were breast-feeding their infants

6 months after leaving the hospital Several weeks

of breast-feeding assures that the mother’s antibod-ies will be present in the infant

Cow’s milk formula resembles its source in terms of type of milk protein, total fat, and calcium

to phosphorus ratio It has been adjusted so that the total protein content, carbohydrate, fat, major minerals, linoleic acid, and vitamins are similar to breast milk The La Leche League International does not recommend substituting formula or breast milk with cow’s milk until the baby is a year old or older (eating the equivalent of three baby food jars

of solid food per day) Unprocessed cow’s milk is not a suitable food for infants for many reasons Cow’s milk contains three times as much protein

as human milk, and this protein is more difficult for babies to digest Manufacturers either presoften

or predigest this protein, or they add whey to

362 indigestion

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adjust the protein ratio Butterfat is also poorly

digested by infants; therefore it is replaced by

veg-etable oils Because the higher concentration of

phosphate and other dissolved minerals in cow’s

milk increases the burden on immature kidneys,

minerals are adjusted to resemble breast milk

Lac-tose or corn syrup solids are used to adjust the

car-bohydrate content Bovine milk protein contains

much more of the essential amino acid

phenylala-nine than human milk protein This situation could

affect infants who cannot tolerate high

phenylala-nine for genetic reasons (see PHENYLKETONURIA)

Cow’s milk in infant formulas sometimes

trig-gers an ALLERGY, especially if there is a family

his-tory of allergies Cow’s milk-based formula as a

supplement to breast-feeding is less of a problem

when the baby is six months or older

For infants who are sensitive to cow’s milk,

liq-uid formulas containing soy protein fortified with

the essential amino acid methionine and with

soy-bean oil are available A variety of formulas are

prepared from coconut oil and corn oil, but these

oils contain very little alpha linolenic acid, an

essential fatty acid Human milk contains

substan-tial amounts of a large fatty acid called DOCOSA

-HEXAENOIC ACID (DHA) DHA, which is necessary

for normal brain and eye development, is not

added to formula There is a consensus that

for-mula should at least contain linolenic acid, the

pre-cursor of DHA, which the infant’s body may

convert to DHA

A wide variety of infant formulas is available to

meet special needs Infants with lactose intolerance

can drink formulas in which lactose is replaced by

other carbohydrates Formulas can be adapted to

adjust protein ratio, linoleic acid content, or to

lower sodium content Special formulas are

avail-able for preterm babies

Ready-to-use formula, as well as powdered

formula, sometimes contains ALUMINUM This is

not a problem for babies with normal kidneys;

however, premature babies may tolerate it poorly

CARRAGEENAN-containing formula should not be

given to premature infants This SEAWEEDproduct is

used to stabilize fat by forming gels in milk

In the past, the infant formula industry

employed questionable marketing practices in

developing countries, which led to a 1977

con-sumer boycott against the Swiss-based Nestlé

com-pany For example, they dressed staff in hospital garb while introducing infant formula to new mothers, and used misleading ads In 1979, Nestlé, which accounted for 50 percent of formula sales to the developing world, and the U.S government formally agreed to voluntary guidelines that banned marketing abuses in developing nations

In 1981, the United Nations World Health Orga-nization voted overwhelmingly to approve an international code of conduct to restrict advertising and marketing of baby formula, which can lead to infant malnutrition and death when improperly used Although not binding, the new guidelines apply to infant formula promotion in industrialized nations as well as developing nations Proper use of infant formula is often impossible in poorer areas

of the world, where the water used to mix the for-mula is often contaminated (See also BABY FOOD.) Ryan, A S “The Resurgence of Breast-feeding in the

United States,” Pediatrics 99, no 4 (1997): e12.

Scariati, Paula D “Risk of Diarrhea Related to Iron Con-tent of Infant Formula: Lack of Evidence to Support the Use of Low-Iron Formula as a Supplement for

Breast-fed Infants,” Pediatrics 99, no 3 (1997): e2.

inflammation A defensive response by the body

to irritation, injury, or infection usually character-ized by heat, redness, swelling, and pain at the injury site This response is triggered by physical agents, chemical agents, or disease-producing organisms Swelling is due to increased blood ves-sel leakage of fluids, and redness is due to the increase in diameter of blood vessels, (especially capillaries) so that they carry more blood With increased vessel leakage, substances normally retained in blood such as water, antibodies, phago-cytic cells, and clot-forming components, migrate into tissues at the site of injury

Cellular Materials that Promote Inflammation

HISTAMINE, kinins, PROSTAGLANDINS, LEUKOTRIENES, and complement contribute to inflammation Hist-amine, derived from the amino acid HISTIDINE, is released from white cells (basophils), mast cells, and other cells when injured Kinins are proteins that induce vasodilation, increase vessel leakiness, attract phagocytic cells, and cause pain Prosta-glandins are hormone-like materials that function

inflammation 363

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in the immediate area where they are produced.

They play many roles, including intensifying pain

and promoting fever, which helps combat

infec-tions Leukotrienes are extremely powerful

inflam-matory agents Complement is a group of blood

proteins that stimulate histamine release, destroy

bacteria, and promote phagocytosis (engulfing

other cells and fragments) Pain can result from

injured nerves or from irritation by released

micro-bial products Inflammation generates free radicals,

which are highly damaging chemical fragments

Chronic inflammation therefore can produce

cellu-lar damage and oxidative stress, leading to an

unbalanced immune response Many chronic

degenerative diseases involve inflammation and

oxidative damage Examples include rheumatoid

arthritis, atherosclerosis, as well as side effects

resulting from radiation and chemotherapy during

cancer treatment

Nonsteroidal anti-inflammatory drugs (NSAIDS)

are often used to combat inflammation These

drugs, which are often nonprescription items, can

themselves cause damage to the stomach or

intesti-nal lining; in some instances they may harm the

liver when used in excess (See also ASPIRIN;

IMMUNE SYSTEM.)

inflammatory bowel disease (IBD) A chronic

inflammation of the intestinal wall involving

painful swelling and open sores Eventually, the

intestinal wall becomes scarred, which narrows

the intestinal opening IBD affects 1 million to 2

million Americans It differs from CELIAC DISEASE, a

grain (especially wheat) intolerance, and from IRRI

-TABLE BOWEL SYNDROME (spastic colon), a much

more common, less serious condition involving

muscle contractions, rather than chronic

inflam-mation Two distinct disorders are classified as

inflammatory bowel disease: CROHN’S DISEASEand

ulcerative COLITIS Crack-like ulcers and abnormal

granular growths in the intestine often accompany

Crohn’s disease, while ulcerative colitis occurs only

in the large intestine and involves inflammation

and ulceration

IBD symptoms include persistent (sometimes

bloody) DIARRHEA, flatulence, cramps, low-grade

fever, and weight loss, as well as problems such as

ARTHRITISand inflamed eyes or skin Children may

be affected by retarded growth and retarded

sex-ual development IBD increases the risk of colon

CANCER IBD is reported mainly in developed countries, where it is most common between the ages of 12 and 40 The causes of IBD are unknown It does not seem to be caused by stress One theory is that IBD is an AUTOIMMUNE DISEASE in which the per-son’s own immune system attacks the intestine Clearly, immune imbalances seem to play a part Another view is that bacteria, viruses, or toxic chemicals initiate IBD Food sensitivity has also been implicated in some cases

Conventional medical treatment involves drugs and/or surgery Drugs reduce inflammation and can lead to a remission However, steroids have side effects like high blood pressure, diabetes, and thinning of bones Typical treatment recommenda-tions include:

• Eating a well-balanced diet that provides ade-quate nutrients to maintain and repair the intestinal tract It is important to correct any nutrient deficiencies caused by the disease (especially iron, folate, and calcium)

• Eating several small meals throughout the day This may be more effective for good digestion and assimilation than eating three big meals

• Avoiding irritating foods that could increase inflammation These differ from person to per-son, though seeds, nuts and corn, lactose and dairy products, fried or greasy foods, and coffee are often the culprits

• Getting enough exercise and managing stress Severe stress suppresses the immune system and encourages inflammation

• Seeking expert medical advice The National Foundation for Ileitis and Colitis provides important information to patients and their families

Gross, V et al “Free Radicals in Inflammatory Bowel Dis-eases—Pathophysiology and Therapeutic

Implica-tions,” Hepato-Gastroenterology 41 (1994): 320–327.

inhibition Restricting the activity of a cellular or physiologic process Several different mechanisms are involved in inhibition

Hormones Hormones may serve as inhibiting agents because they can inhibit the release of other

364 inflammatory bowel disease

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hormones Thus, the ovaries produce inhibin (a

hormone that inhibits the secretion of the ovarian

hormones), follicle stimulating hormone, and

lut-enizing hormone, at the end of the menstrual

cycle Elevated CORTISOL from the adrenal glands

inhibits the release of ADRENOCORTICOTROPIN

(ACTH) from the PITUITARY GLAND ACTH stimulates

the release of cortisol

Enzymes Enzymes are protein catalysts that

lose activity when blocked by inhibitors

Competi-tive enzyme inhibitors are compounds that mimic

the chemical that the enzyme usually alters

Enzyme poisons like toxic heavy metals—LEAD,

MERCURY, and CADMIUM—can bind to enzymes

without competing with substances and can

per-manently inactivate enzymes

Certain key regulatory enzymes may be

re-versibly inhibited by the accumulation of key

prod-ucts of metabolism One example is the feedback

inhibition of a METABOLIC PATHWAY, a sequence of

functionally linked enzymes In this type of

inhibi-tion, a surplus of the final product of the pathway

can inhibit the pathway For example, the buildup

of ATP, the energy currency of the cell, can inhibit

enzyme systems like GLYCOLYSISthat generate ATP

Such feedback mechanisms help the cell avoid

wasteful overproduction of products (See also

ENDOCRINE SYSTEM; INDUCTION.)

inosine One of the basic compounds composing

cells and a precursor to adenosine, an important

energy molecule and building block of DNA and

RNA Although some European scientists believed

it could have energy-boosting effects, controlled

studies concluded that inosine does not improve

athletic performance Athletes often take between

5,000 and 6,000 mg of inosine a day, but research

does not support the use of this supplement in any

amount

However, some animal research studies have

suggested it may be helpful in the treatment of

stroke and other central nervous system disorders

Inosine occurs in organ meats and brewer’s yeast

and can be taken as a supplement Although there

are no reports of side effects, any inosine that is not

used by the body is converted to uric acid, which

could be a problem for people at high risk for gout

Safety data are inadequate for pregnant and

breast-feeding women

Starling, R D., T A Trappe, K R Short et al “Effect of Inosine Supplementation on Aerobic and Anaerobic

Cycling Performance,” Medicine and Science in Sports

and Exercise 28 (1996): 1,193–1,198.

inositol (myoinositol) An essential building block of cell membrane LIPIDS Chemically, inositol

is a cyclic ALCOHOLwith six hydroxyl groups, one per carbon atom Inositol is a constituent of phos-phatidylinositol, a component of inner-cell mem-branes Derivatives of inositol function as HORMONE

relay signals in cells Diverse hormones such as

VASOPRESSIN(from the pituitary gland), EPINEPHRINE

(from the adrenal gland) and releasing factors from the HYPOTHALAMUSstimulate the release of inositol triphosphate from phosphatidylinositol

Animal studies show that inositol may protect against ATHEROSCLEROSISand against hair loss Inos-itol is also supposed to help reverse nerve damage caused by diabetes in animals Oral supplementa-tion in human diabetics has not verified this result Diabetics should consult their health care providers before taking inositol supplements Inositol has a low toxicity

Nutritionists have not yet established the opti-mum amount of inositol in the diet It is widely dis-tributed in food and is also manufactured in the body Sources include CITRUS FRUIT (except lemons), CANTALOUPE, whole grain bread, cooked beans, green beans, and nuts Inositol occurs in grains such as PHYTIC ACID, in which six phosphate groups are attached to the inositol molecule Phytic acid can bind minerals and limit their uptake Safety data are inadequate for pregnant and breast-feeding women (See also VITAMIN.)

Shamsuddin, Abulkalaman M “Inositol Phosphates Have

Novel Anti-Cancer Function,” Journal of Nutrition 125,

supp 3 (1995): 725S–732S.

insulin A protein HORMONE, secreted by beta cells

in the PANCREAS, that stimulates the uptake of

BLOOD SUGAR by many tissues Insulin counteracts the effects of GLUCAGON, the pancreatic hormone responsible for raising blood sugar Insulin is used therapeutically to treat DIABETES MELLITUS and is either purified from pork or beef pancreas or is ge-netically engineered of human origin Insulin is produced by small cell clusters in the pancreas

insulin 365

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