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Tiêu đề Biodegradable Gellan Gum Hydrogels Loaded with Paclitaxel for HER2+ Breast Cancer Local Therapy
Tác giả Celia Nieto, Milena A. Vega, Víctor Rodríguez, Patricia Pérez-Esteban, Eva M. Martín del Valle
Trường học University of Salamanca
Chuyên ngành Chemical Engineering
Thể loại Research Article
Năm xuất bản 2022
Thành phố Salamanca
Định dạng
Số trang 15
Dung lượng 9,62 MB

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Nội dung

Hydrogels loaded with chemotherapeutics are promising tools for local tumor treatment. In this work, redoxresponsive implantable hydrogels based on gellan gum were prepared as paclitaxel carriers for HER2-positive breast cancer therapy. To achieve different degrees of chemical crosslinking, hydrogels were synthesized in both acetate buffer and phosphate buffer and crosslinked with different concentrations of L-cysteine.

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Available online 15 June 2022

0144-8617/© 2022 The Authors Published by Elsevier Ltd This is an open access article under the CC BY-NC-ND license ( http://creativecommons.org/licenses/by-nc-nd/4.0/)

Biodegradable gellan gum hydrogels loaded with paclitaxel for HER2+

breast cancer local therapy

Celia Nietoa, Milena A Vegaa, Víctor Rodrígueza, Patricia P´erez-Estebanb, Eva M Martín del

Vallea,*

aChemical Engineering Department, Faculty of Chemical Sciences, University of Salamanca, Salamanca 37008, Spain

bCollege of Health and Life Sciences, School of Biosciences, Aston University, Birmingham B4 7ET, UK

A R T I C L E I N F O

Keywords:

Gellan gum

Hydrogel

Local chemotherapy

HER2-positive breast cancer

Paclitaxel

β-Cyclodextrin

Glutathione

A B S T R A C T Hydrogels loaded with chemotherapeutics are promising tools for local tumor treatment In this work, redox- responsive implantable hydrogels based on gellan gum were prepared as paclitaxel carriers for HER2-positive breast cancer therapy To achieve different degrees of chemical crosslinking, hydrogels were synthesized in both acetate buffer and phosphate buffer and crosslinked with different concentrations of L-cysteine It was shown that both, the type of buffer and the L-cysteine concentration used, conditioned the dynamic modulus, equilibrium swelling rate, porosity, and thermal stability of the hydrogels Then, the biocompatibility of the hydrogels with the most suitable porosity for drug delivery applications was assessed Once confirmed, these hydrogels were loaded with paclitaxel:β-cyclodextrin inclusion complexes, and they showed a glutathione-

responsive controlled release of the taxane Moreover, when tested in vitro, paclitaxel-loaded hydrogels

exhibi-ted great antitumor activity Thus, they could act as excellent local tailored carriers of paclitaxel for future, post- surgical treatment of HER2-overexpressing breast tumors

1 Introduction

Breast cancer is currently considered as one of the diseases with the

highest mortality rate in woman worldwide (Tang et al., 2021), with

685,000 deaths associated with female breast cancer being reported last

year alone (Sung et al., 2021) Among the different alternatives that

exist for its treatment, surgical resection is the gold standard clinical

strategy (Bu et al., 2019; Tang et al., 2021; Zhuang et al., 2020)

Nevertheless, despite much improvement in surgical techniques,

effi-cient inhibition of breast cancer recurrence still presents a challenge

The main reason for this is that residual tumor cells can remain in

sur-gical margins (Askari et al., 2020; Bastiancich et al., 2017), particularly

in patients who have undergone breast-conserving therapy (Qu et al.,

2015)

To reduce the incidence of relapse, radiotherapy and chemotherapy

are routinely administered in the clinical setting after tumor resection

However, both treatments are associated with high toxicity and severe

systemic side effects (Bu et al., 2019; Tang et al., 2021) In addition,

since these forms of treatment must begin in the weeks following surgery

to allow the patient's health to recover, residual infiltrative cancer cells

can keep proliferating in the meantime (Bastiancich et al., 2017; Bu

et al., 2019; Zhuang et al., 2020) Moreover, resistance to chemotherapy may be promoted, in addition to other factors such as hypoxia or al-terations in the signaling pathways of cancer cells, by the limited tar-getability of the anticancer drugs (Askari et al., 2020; Kibria & Hatakeyama, 2014) For these reasons, local delivery of chemothera-peutics in the tumor resection cavity is becoming increasingly desirable for breast cancer treatment (Tang et al., 2021) Compared to systemic therapies, local chemotherapy can prevent drugs from being non- specifically distributed and can avoid off-target toxicities Moreover, local chemotherapy may eliminate the latency time of post-surgical systemic chemotherapy (Askari et al., 2020; Tang et al., 2021; Zhuang

et al., 2020)

Among the different types of drug delivery systems (DDS) designed for antitumor local therapies, hydrogels are, in particular, generating greater interest, as their mechanical properties can be tailored to mimic those of the extracellular matrix (ECM) of living tissues (Askari et al.,

2020) Furthermore, most of these three-dimensional hydrophilic net-works are made from natural polymers; thus, they are biocompatible, biodegradable and easily modifiable, in addition to having high drug-

* Corresponding author

E-mail address: emvalle@usal.es (E.M Martín del Valle)

Contents lists available at ScienceDirect Carbohydrate Polymers

journal homepage: www.elsevier.com/locate/carbpol

https://doi.org/10.1016/j.carbpol.2022.119732

Received 3 February 2022; Received in revised form 30 May 2022; Accepted 9 June 2022

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loading capacities (Abasalizadeh et al., 2020; Darge et al., 2019; Misra &

Acharya, 2021; Sharma & Tiwari, 2020) Among the most common

natural polymers, gellan gum (GG) is gaining attractiveness for

biomedical purposes, as it is stable and has appropriate mechanical

properties, acid and heat resistance and inotropic sensitivity GG is an

extracellular polysaccharide, which contains repeating units of β-D-

glucose, L-rhamnose, and D-glucuronic acid in a 2:1:1 M ratio (Das &

Giri, 2020; Palumbo et al., 2020), that can undergo thermally reversible

gelation after a coil-helix transition in the presence of mono- (K+, Na+)

or divalent (Ca2+) cations (Bacelar et al., 2016; Prajapati et al., 2013;

Soleimani et al., 2021) Similarly, GG can be chemically crosslinked to

maintain stable biomaterial structures for longer periods Previous

works involving this polysaccharide have been reported regarding its

use for the delivery of several anticancer drugs (paclitaxel, doxorubicin,

erlotinib and clioquinol, among others) to improve their solubility,

intra-tumoral specificity, and drug release profile via hydrogels, patches

and nanoconfigurations (Villareal-Otalvaro & Coburn, 2021) In the

specific case of paclitaxel (PTX), GG has been employed to develop in

situ-gelling liposome-in-gel composites containing this drug for local

bladder cancer treatment, and nanohydrogels delivering the taxane

along with prednisolone for prostate cancer and inflammatory

carci-noma applications (D'Arrigo et al., 2014; GuhaSarkar et al., 2017)

However, GG has not yet been used to fabricate PTX-loaded implantable

hydrogel patches for local, stimuli-responsive treatment of HER2-

positive (HER2+) breast tumors Therefore, the main aim pursued in

this work was to develop, characterize and validate in vitro PTX-

releasing GG hydrogel patches that would be suitable for this novel

application: local and redox-responsive antitumor therapy of HER2+

breast tumors

Consequently, GG hydrogels (HGGs) were prepared in two solutions

with different pH and ionic compositions (acetate buffer [AB] vs

phosphate buffered saline [PBS]) and were disulfide-crosslinked with

different L-cysteine (L-Cys) concentrations utilizing the carbodiimide chemistry to improve their stability while achieving responsiveness to external reducing stimuli, such as the high glutathione (GSH) concen-trations existing in malignant breast cells (Li et al., 2020; P´erez et al.,

2014) The main aim of synthesizing HGGs in different buffers and with different L-Cys concentrations was examining how these parameters conditioned their crosslinking degree and, therefore, their dynamic modulus, equilibrium swelling rate, porosity, and thermal stability Then, all these hydrogel properties were analyzed and, based on the results obtained, those HGGs with the most appropriate characteristics for drug delivery applications were selected to be loaded with PTX This taxane was previously included in β-cyclodextrin (βCD) molecules to improve its limited aqueous solubility (Nieto et al., 2019; Tian et al.,

2020), and the resulting complexes (PTX:βCDs) were included in the GG patches to enhance the redox-controlled release of PTX while trying to improve its bioavailability and off-target toxicity through a potential local application (Scheme 1) Antitumor activity of the HGGs loaded

with the PTX:βCD complexes was evaluated in vitro after analyzing their

biocompatibility, and the results obtained showed that they may be a promising strategy for post-surgical chemotherapy of HER2- overexpressing breast tumors with elevated GSH intracellular concentrations

2 Materials and methods

2.1 Materials

Gelzan™ CM (G1910, average molecular weight: 1000 kg/mol; low- acyl [0.2 %]; monosaccharide composition: β-D-glucose:L-rhamnose:D- glucuronic acid [2:1:1]), β-cyclodextrin (βCD, minimum 98 %),

pacli-taxel (PTX, from semisynthetic, >97 %), L-cysteine (L-Cys, 97 %), lyso-zyme human, 1-ethyl-3-(3-dimethylaminopropyl) carbodiimide (EDC),

Scheme 1 Schematic representation of the preparation of the HGG patches, chemically crosslinked with different concentrations of L-Cys and loaded with PTX:β

CD complexes

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N-hydroxy succinimide (NHS), thiazolyl blue tetrazolium bromide

(MTT), phosphate buffered saline (PBS, powder [NaCl [137 mM], KCl

[2.7 mM], Na2HPO4 [10 mM], KH2PO4 [1.8 mM], pH 7.4) and L-

Glutathione reduced (>98 %) were all obtained from Sigma Aldrich (St

Louis, MO, USA) Dimethyl sulfoxide (DMSO, >99 %) and Corning™

penicillin/streptomycin solution (100×: penicillin [100 UI/ml] and

streptomycin [10,000 μg/ml]) were purchased from Thermo Fisher

Scientific (Waltham, MA, USA) Calcein AM and propidium iodide (PI,

Ready Probes™) were obtained from Invitrogen (Carlsbad, CA, USA)

Potassium bromide (for IR), acetic acid glacial, citric acid anhydrous,

sodium acetate anhydrous, sodium citrate, sodium chloride, tris

hy-drochloride and absolute pure ethanol (EtOH) were all obtained from

Panreac AppliChem (Castellar del Vall`es, Barcelona, Spain) Dubelcco's

Modified Eagle's Medium (DMEM) and fetal bovine serum (FBS,

quali-fied, HI) were purchased from Gibco (Gaithersburg, MD, USA) Finally,

lactate dehydrogenase activity colorimetric assay kit (product code:

ab102526) was obtained from Abcam (Cambridge, UK)

2.2 Synthesis of HGG patches

To prepare the HGG patches, Gelrite® (Gelzan™) was chosen among

the main different commercial forms of GG because it disperses and

hydrates well in deionized water (H2O[d]) and is inert to most biological

growth media additives (Prajapati et al., 2013) In this way, Gelzan™

was dissolved (1.5 % [w/v]) both in 80 ◦C AB (0.05 M, pH 4.0) and in

80 ◦C PBS (pH 7.4) (Matricardi et al., 2009; Oliveira et al., 2016) Once

homogeneous solutions were obtained, the temperature was lowered to

50 ◦C Solutions of EDC (2.9 mg/ml) and NHS (4.8 mg/ml) were later

incorporated consecutively (1:50 [v/v]) After stirring briefly, L-Cys

solutions of different concentrations (1.5, 3, and 4.5 mg/ml) were added

(1:50 [v/v]) to achieve different degrees of GG chemical crosslinking

(Wu et al., 2018; Yu et al., 2020) Final solutions were poured into dishes

and left for gelation at room temperature overnight

2.3 Rheology

Rheological measurements of 2-mm-thick HGGs were performed

using an AR 1500 Ex rheometer (Waters Corporation, Milford, MA, USA)

equipped with an aluminum parallel plate geometry (plate diameter 40

mm, gap distance 1 mm) HGG samples were prepared using 33 mm-

diameter dishes as templates, carefully unmolded preventing breakage

and placed on the lower plate of the rheometer To evaluate their

stiff-ness, dynamic oscillation-frequency tests were carried out in duplicate

in the 0.01–10 Hz range at 25 ◦C and 37 ◦C by applying a γ = 0.01

constant deformation in the linear viscoelastic region This region was

preliminary assessed using stress sweep tests (Matricardi et al., 2009)

(data not shown)

2.4 Swelling test

The swelling ability of the different HGGs was assessed via a general

gravimetric method Variations in weight were recorded over time when

the HGGs were soaked in solutions of different pH and ionic strength:

H2O(d) (purified with the Economatic Wasserlab equipment [Barbat´ain,

Navarra, Spain]); commercial mineralized water (H2O[c]); NaCl

solu-tion (0.015 M); tris buffer (0.05 M); citrate buffer (0.1 M); AB (0.04 M);

PBS (1×); and DMEM supplemented with FBS and antibiotics Briefly,

after gelation, hydrogel disks (35 mm diameter, 8 mm height) were

frozen at − 80 ◦C, lyophilized overnight (LyoQuest lyophilizer, Telstar,

Lisbon, Portugal), and weighed Then, hydrogels were immersed in the

previously mentioned solutions (50 ml), removed after different time

points, wiped superficially with bibulous paper, weighed again, and

introduced in the same solutions (Coutinho et al., 2010; Li et al., 2021;

Morello et al., 2021) The swelling ratios at time t (Qt) and when HGGs

reached equilibrium (Q∞) were defined according to Eqs (1) and (2),

respectively, where m0 is the initial weight of the dried gels (g), mt is the

weight of the swelled gels after time t (g), and m∞ is the weight of the swelled gels at the equilibrium (g) (Schott, 1992)

Q t=m tm0

Q∞=m∞− m0

To better describe the swelling behavior of the HGGs, a swelling kinetic study was performed at the initial stage of swelling, until hydrogels reached equilibrium For this purpose, Eqs (1) and (2) were adjusted to a pseudo-second-order kinetic model, as described by Schott

in 1992 (Supplementary Material) It was considered that homogeneous uptake of the solutions occurred throughout the hydrogel polymer networks

2.5 Evaluation of the crosslinking density

The effective crosslinking density (dx, mol/ml) of the six different prepared HGGs was determined according to Eq (3):

d x= 1

where ϑ is the specific volume of the polymer (ml/g) and Mc is the average molecular mass between crosslinkings (g/mol), which was determined by the Flory-Rehner equation (Eq (4)):

M c= ρ p V s V 1/3

r

[

ln(1 − V r) +V r+XV2

r

where V s is the molar volume of the solvent (ml/mol), ρ p is the density of the polymer (g/ml), X is the parameter of interaction between the sol-vent and the polymer (which has a value of 0.81 ± 0.05 for aqueous solutions of GG (Safronov et al., 2019) and Vr is the polymer volume fraction calculated from Eq (5)

V r=

[

1 +ρ p

ρ s

(

M a

M b

) +ρ p

ρ s

]

(5) where Ma is the swollen hydrogel weight (g), Mb is the weight of the dried hydrogel before the swelling experiment (g) and ρ is the density of the solvent (g/ml) (Afinjuomo et al., 2019; Sabadini et al., 2018)

2.6 Morphological analysis and porosity determination after freeze- drying

The porous structure of the different HGGs synthesized was analyzed

by scanning electron microscopy (SEM) (ESEM Quanta 200 FEG, FEI, Hillsboro, OR, USA) HGG samples were freeze-dried, coated with gold and cross-sectioned Then, samples were imaged at an accelerating voltage of 15 kV 8 to 10 images were acquired from different areas of each sample and the average diameter of the micro- and macropores

existing in the HGGs was determined via image analysis (ImageJ

soft-ware) (Hua et al., 2016; Lee et al., 2020)

In addition, HGG porosity was measured using Archimedes' princi-ple Once synthesized, all hydrogel samples were freeze-dried and completely immersed in tubes filled with absolute EtOH After 24 h, HGGs were removed from the tubes and their porosity was calculated according to Eq (6):

Porosity (%) = W2− W3−W s

W1− W3

where W1 is the weight of the tube filled with EtOH (g), W2 is the weight

of the tube filled with EtOH 24 h after immersion of the freeze-dried HGGs (g), W3 is the weight of the tube filled with EtOH after HGG removal (g) and WS is the weight of the freeze-dried HGGs (g) (Goodarzi

et al., 2019)

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2.7 Fourier transform infrared (FTIR) characterization

The chemical structure of all HGG samples, as well as that of GG, was

analyzed by FTIR spectroscopy (Spectrum Two™ spectrometer, Perkin

Elmer, Waltham, MA, USA) at the wavelength range of 900–4000 cm− 1

and compared Freeze-dried samples were ground to powder, dried at

37 ◦C for 3 days to remove any possible residual water, prepared with

KBr pellets, and scanned

2.8 Thermogravimetric analysis (TGA)

HGG thermal stability was analyzed by TGA (DSC Q100 calorimeter,

Waters Corporation, Milford, MA, USA) and compared to that of GG

alone HGGs were freeze-dried, and all samples were later ground to

powder and heated at a rate of 10 ◦C/min from 50 ◦C to 600 ◦C under a

nitrogen atmosphere to obtain the thermogravimetric (TG) curves

2.9 Compression test

The compressive modulus of cylinder samples (35 mm diameter, 8

mm height) of the HGGs chosen to be later loaded with the PTX:βCD

complexes was determined by spherical indentation testing Thus, a

spherical indenter was employed as plunger (Fig S1), the force-

indentation curve for the samples was recorded, and the effective

stiff-ness of the hydrogels was extracted For this purpose, the indentation

curves obtained were fitted to Hertz's contact model (Eq (7)) (Srivastava

et al., 2017)

F = − 16E2

̅̅̅̅̅̅̅̅

Rd3

where F was the force applied by the indenting bead (N), E2 was the

Young's modulus of the different HGG samples (kN/m2), R was the

diameter of the bead (6 mm) and d was the indentation depth (mm)

HGG average Young's modulus was determined from the slope obtained

after plotting F vs d3/2 Three parallel samples were tested to obtain an

average

2.10 Hydrogel in vitro degradation

The degradation rate of the HGGs (35 mm diameter, 8 mm height)

later loaded with the PTX:βCD complexes was investigated in vitro

through weight loss under simulated tumor extracellular pH conditions

Once weighed (m0, g), HGGs were placed in duplicate in beakers

con-taining lysozyme solution (1 mg/ml in PBS (pH 6.8)) and incubated for

9 days at 37 ◦C under gentle shaking (50 rpm) HGGs were weighed daily

(mt, g) after wiping their surface with bibulous paper, and their weight

loss (mr) was determined according to Eq (8) (Huang et al., 2020; Lu

et al., 2022; Panczyszyn et al., 2021; Xu et al., 2018):

m r(%) =m0− m t

m0

2.11 Cell culture and hydrogel biocompatibility in vitro

Human HER2+ breast carcinoma BT474 cells and stromal HS5 cells

were grown in DMEM supplemented with 10 % (v/v) FBS and 1 % (v/v)

penicillin/streptomycin, and cultured in an atmosphere of 5 % CO2 at

37 ◦C

HGG biocompatibility was doubly assessed by MTT assays and live/

death staining BT474 and HS5 cells were seeded in 24-well plates

(12,000 cells/ml), grown for 24 h for attachment, and cultured with

HGG samples that were allowed to gel for 90 min (23.1 % [v/v],

pre-viously sterilized by UV radiation) Cells were incubated for 72 h and

their survival rate was studied by MTT colorimetry tests At specified

times (including 24, 48 and 72 h), 110 μl MTT solution (5 mg/ml in PBS)

was added to the wells, cells were incubated further for 1 h at 37 ◦C, and

the resulting formazan salts were dissolved in DMSO (500 μl/well) (Rahnama et al., 2021) The optical density (OD) of each well was recorded using a microplate reader (EZ Microplate Reader 2000, Bio-chrom, Cambridge, UK) at a wavelength of 550 nm after shaking for 10 min Cells not exposed to HGG samples were used as a blank control group, and three independent samples were included for each time in-terval and experimental group

BT474 and HS5 cells were also seeded in 8-well glass-bottom slides (12,000 cells/ml), grown for 24 h and exposed or not to HGG samples

(23.1 % [v/v], also sterilized by UV radiation) for a further 24 h Then,

15 min before imaging the cells by confocal laser scanning microscopy (CLSM), calcein AM (1 μg/ml) and PI (5 μg/ml) were used to stain alive (green) and dead (red) cells, respectively (Huan et al., 2022) Samples (two independent ones for each experimental group) were washed with PBS solution before CLSM imaging (TCS SPS, Leica Microsystems, Wetzlar, Germany)

2.12 HGG loading with PTX:βCD complexes

To improve PTX aqueous solubility, PTX:βCD inclusion complexes were obtained following the freeze-drying method described by Alcaro

et al., 2002 Briefly, PTX (1 mg) was dissolved in absolute EtOH (1.2 ml), and βCDs (1.2 mg) were dissolved in H2O(d) (1.4 ml) Next, the βCD solution was added to the PTX solution, and the resulting hydroalcoholic solution was kept under agitation (100 rpm) for 5 h at room temperature and in the dark Later, it was frozen at − 80 ◦C and freeze-dried (Nieto

et al., 2019) The white powder obtained was dissolved in H2O(d), achieving a 0.185 mM PTX working concentration

Subsequently, to load HGG patches with the PTX:βCDs prepared, hydrogel synthesis was performed as described above PTX:βCD solu-tions were added while the gelation process was taking place, once the L- Cys solutions (3 mg/ml) were incorporated (Ning et al., 2020) HGGs loaded with the chemotherapeutic (HGGs@PTX) were allowed to cool in dishes or multi-well plates for their complete gelation

2.13 PTX-release from HGGs in vitro

Once obtained, crosslinked HGGs@PTX were allowed to gel for 90 min and washed with PBS to remove the unloaded taxane before per-forming drug release experiments in duplicate Next, hydrogel patches (35 mm diameter, 8 mm height) were soaked in crystallizing dishes containing slightly acidic PBS (60 ml, pH 6.8) and incubated at 37 ◦C at

40 rpm for 72 h To mimic the intracellular redox potential of tumor cells, GSH was added in high concentrations (10 mM) to the release medium of some HGG samples (P´erez et al., 2014; Robby et al., 2021) At pre-determined times, 0.5 ml aliquots were taken out, and equal vol-umes of acidic PBS (containing or not GSH [10 mM]) were added to maintain a constant volume in the crystallizing dishes The amount of PTX released was calculated by comparing the absorbance of the ali-quots at 230 nm (UV-1800 spectrophotometer, Shimadzu Corporation, Kioto, Japan) with a previously measured calibration curve obtained from a PTX dilution series Aliquots of the release media of non PTX- loaded HGGs were used as a blank Cumulative PTX release (%) from the different HGGs samples was determined according to Eq (9) and plotted against time (Fang et al., 2021; Rezk et al., 2019; Vu et al.,

2022)

PTX released (%) = Total PTX released

Moreover, PTX release kinetics were studied through four different

mathematical models, i.e., zero-order, first-order, Korsmeyer-Peppas

and Higuchi models A description of the method is reported in the Supplementary Material

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2.14 Antitumor activity of HGGs@PTX in vitro

HGG@PTX antitumor activity was analyzed in vitro on two different

human HER2-overexpressing breast carcinoma cell lines: BT474 and

SKBR3 (Nieto et al., 2019)

Cells were cultured as previously indicated, and MTT assays and

live/death staining were conducted following the same protocols as

before to doubly assess crosslinked HGG@PTX cytotoxicity

Neverthe-less, this time, BT474 and SKBR3 cells were exposed to HGGs (23.1 %

[v/v]), HGGs@PTX (23.1 % [v/v]) and PTX:βCDs (in an equivalent

concentration to that loaded to the HGGs (30.8 μM)) Besides, live/death

staining was performed 48 and 72 h after cell exposure to the different

treatment conditions Again, cells not exposed to HGG samples served as

a blank control group in both assays

In addition, lactate dehydrogenase (LDH) leakage assays were

car-ried out according to LDH activity detection kit manufacturer's

in-structions to analyze BT474 and SKBR3 membrane damage after

treatment with the HGGs[3LCys]@PTX for 48 h Group distributions

and PTX:βCDs and HGG[3LCys]@PTX concentrations similar to those in

the MTT assays were employed The absorbance of the LDH expression

was assessed at 450 nm using a microplate reader

2.15 Statistical analysis

All data were reported as mean ± standard deviation (SD) Specific

comparison between groups was carried out with unpaired Student's t-

tests, while one-way ANOVA was used for multiple-group comparison

p-values <0.05 were considered to be statistically significant When

statistically significant differences were found when performing one-

way ANOVA, Tukey test was carried out as post-hoc analysis

3 Results and discussion

3.1 Preparation of HGGs with different degrees of chemical crosslinking

One of the characteristics of GG that has led to its increased use for biomedical purposes is its ionotropic sensitivity (Das & Giri, 2020;

Palumbo et al., 2020) In this way, obtaining HGGs is possible because, when mono- or divalent cations are present in a solution, GG can un-dergo thermally reversible gelation after transition from a coiled form at

high temperature (>80 ◦C) to a double-helix structure when cooled (Bacelar et al., 2016; Prajapati et al., 2013) Thus, HGG consistency can

be modified, apart from altering the concentration of the gum, by adding different ions to GG solutions (Das & Giri, 2020; Palumbo et al., 2020)

Fig 1 Frequency sweeps of the different synthesized HGGsAB and HGGsPBS performed at 25 ◦C (A–C) and 37 C (B–D) Filled symbols represent G′values, while empty symbols are G′′values The concentration of L-Cys used to crosslink the different hydrogels is indicated between brackets (in mg/ml)

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For this reason, as indicated in Scheme 1, two buffers of different ionic

composition and pH (AB vs PBS) were used in this work to synthesize

HGGs with the aim of analyzing how they conditioned the

physico-chemical properties of the hydrogels obtained (HGGsAB vs HGGsPBS

respectively) (Matricardi et al., 2009; Oliveira et al., 2016) In addition,

to enhance their stability and make them redox-responsive, HGGs were

chemically crosslinked with L-Cys (Du et al., 2012), which was employed

in three different concentrations (1.5, 3, and 4.5 mg/ml) to later choose

the most suitable hydrogels to act as PTX delivery systems EDC

chem-istry was used to carry out the crosslinking because, unlike other

com-pounds frequently used to prepare chemical hydrogels, EDC and NHS

are not cytotoxic in concentrations below 0.5 M (Hua et al., 2016;

Panczyszyn et al., 2021) In addition, these compounds have already

been used in the literature to crosslink hydrogels made up of other

polymers (Goodarzi et al., 2019; Pacelli et al., 2018; Výborný et al.,

2019), and the N-hydroxysuccinimidyl ester coupling chemistry is one

of the few conjugation strategies utilized in the development of FDA-

approved protein conjugates (Kang et al., 2021; Pelegri-O'Day et al.,

2014)

3.2 Rheological properties of the different HGGs

Once obtained, the viscoelastic properties of the six different

syn-thesized types of HGG were determined employing dynamic oscillatory

frequency sweep assays and compared Mechanical spectra recorded

both at 25 ◦C and 37 ◦C can be found in Fig 1

As can be observed in Fig 1, the frequency sweeps obtained

indi-cated that all samples had characteristic gel behavior, since the storage

modulus (G′) was at least 10 times higher than the loss modulus (G′′) in

all cases Moreover, both G′ and G′′ were almost independent of the

frequency, which is a distinctive fact of entangled gels (Matricardi et al.,

2009; Richa & Choudhury, 2019) However, when comparing the

spectra of the different HGGsAB (Fig 1[A–B]) with those of the HGGsPBS

(Fig 1[C–D]), it was observed that G′ values were greater when

hydrogels were prepared in PBS than in AB In this way, HGGsPBS gelled faster and were more viscous than HGGsAB This result was logical considering that PBS contains K+cations and higher concentrations of

Na+cations (>10 times greater) than AB (Table S1) and, therefore, that

it could contribute to achieving greater degree of GG crosslinking

As expected, when the L-Cys content of both HGGsAB and HGGsPBS

was higher, G′values increased due to the existence of more chemical crosslinkings and the consequent formation of stronger 3D networks This trend could be also seen when increasing the measurement tem-perature from 25 ◦C to 37 ◦C, although this increase in temperature resulted in diminished G′values, which were 40–60 % lower than those recorded at 25 ◦C (Matricardi et al., 2009) Hence, this reduction in the elastic modulus suggested that HGG equilibrium constants were thermal sensitive, and that this sensitivity could be related to the initial degree of crosslinking of the HGGs, since G′reduction was less noticeable when hydrogels were disulfide-crosslinked with higher concentrations of L-Cys and when they were synthesized in PBS instead of in AB (Roberts et al.,

2007)

3.3 Swelling behavior of the different HGGs as a function of the medium

pH and ionic strength

Since the rate and degree of swelling of hydrogels are the most important parameters when controlling the release of the drugs with which they may be loaded (Ganji et al., 2010), the swelling kinetics of all HGGs prepared were analyzed as a function of the medium pH and ionic strength (μ) For this purpose, HGG samples were soaked in H2O(d) and

H2O(c) to determine whether their different ionic composition condi-tioned hydrogel swelling capacity Likewise, HGGs were soaked in NaCl solutions, PBS and supplemented DMEM because these media with different ionic strength mimic physiological fluids Moreover, tris buffer, citrate buffer and AB were also employed to perform swelling assays to try to determine how the medium acidity or basicity could condition HGG absorption capacity The properties of all these media can be found

Fig 2 Swelling kinetics of the different HGGsAB (A–C) and HGGsPBS (D–F) as a function of the swelling time when soaked in solutions with different pH and ionic strength at 25 ◦C

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in Table S2

The swelling kinetics obtained for the HGGsAB crosslinked with

different concentrations of L-Cys are shown in Fig 2(A–C), while those of

the three different HGGsPBS can be seen in Fig 2(D–F)

As shown in Fig 2, most HGG samples reached equilibrium after 240

min Thereby, after soaking HGGs in the different media for about 4 h,

there was a balance between the osmotic forces caused by the solutions

when entering the hydrogel macromolecular networks and the cohesive-

elastic forces exerted by the GG chains, which opposed the expansion

For this reason, the experimental data obtained up to 240 min were

adjusted to a pseudo-second-order kinetic model to determine Q∞ and

K∞ values for all HGGsAB and HGGsPBS in the different media (Panpinit

et al., 2020; Schott, 1992) The values obtained for these parameters,

which refer to the theoretical equilibrium swelling capacity and the

swelling rate constant of the HGGs, respectively, are indicated in

Table S3 and S4

When comparing the parameters of the swelling kinetics of both

types of HGGs as a function of their crosslinking degree, it was noticed

that, in general, the greater crosslinking, the lower the HGG swelling

capacity This fact was in line with what was expected since by

increasing L-Cys concentration during the synthesis process, it was likely

that HGG pore size would be reduced, and that hydrogels would take up

less volume when soaked in the different media (Coutinho et al., 2010)

In the same way, as the degree of crosslinking of the HGGsAB was

lower than that for the HGGsPBS, they showed greater swelling capacity

and, therefore, higher Q∞ and K∞ values, especially in the most alkaline

media: H2O(d), H2O(c), tris buffer and DMEM Possibly, as described in

the literature, H+cations could interact with GG negative charges after

penetrating the hydrogel structure, causing greater aggregation of GG

chains at low pH values By contrast, in basic media, OH− anions may

accelerate the electrostatic repulsion of GG chains, causing hydrogels to

experience a hydrolysis-induced swelling behavior and to have higher

swelling rates than in acidic solutions (Cassanelli et al., 2018; De Souza

et al., 2016; Moritaka et al., 1995; Zhou & Jin, 2020) In fact, when

HGGs were soaked in H2O(d) and, especially, in tris buffer, they started

to break after 30 min, possibly because the electrostatic repulsion be-tween the COO− anions was too strong and hydrogels lost their network structure In addition, as shown in Fig 2, the less crosslinked HGGsAB experienced over-swelling when soaked in tris buffer, followed by a deswelling process that took place until they reached equilibrium Probably, since these HGGs could oppose less resistance to the entry of tris buffer in their structure, this phenomenon could take place because

of the difference in osmotic pressure that occurred at the initial stage of the swelling process (Li et al., 2021)

Finally, regarding the effect of the ionic strength of the media on HGG swelling behavior, another phenomenon already described in the literature could be observed: in those media with greater ionic strength (DMEM, PBS, citrate buffer, AB and NaCl solution), HGG swelling occurred in a lesser extent than in media with less ions (H2O[d] and H2O [c]) due to GG ionotropic sensitivity Thus, like H+, cations existing in the solutions in which hydrogels were soaked could interact with GG chains, promoting their aggregation and, therefore, lowering HGG me-dium uptake capacity (Coutinho et al., 2010; Moritaka et al., 1995)

3.4 Crosslinking density of the different HGGs

Besides, since crosslinking density (dx) and average molecular weight between crosslinks (Mc) determine hydrogel swelling capacity and, therefore, hydrogel drug release patterns, dx and Mc of the different HGGs were also determined based on the data obtained in the swelling tests once HGGs reach equilibrium in H2O(d) The values calculated for these parameters, as well as for the different polymer volume fractions (Vr), are reported in Table S5

As can be seen in the Supplementary Material, when greater con-centrations of L-Cys were employed for HGG preparation, the average polymer volume fraction and molecular weight between crosslinkings diminished By contrast and as expected, HGG crosslinking density increased In this way, when greater amounts of crosslinker were

Fig 3 Morphological analysis under SEM of (A) HGGAB[1.5LCys], (B) HGGAB[3LCys], (C) HGGAB[4.5LCys], (D) HGGPBS[1.5LCys], (E) HGGPBS[3LCys] and (F) HGGPBS[4.5LCys] samples

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incorporated, the space for solvent accommodation between GG chains

could be reduced, being this fact in agreement with the results

previ-ously obtained in the swelling tests

3.5 Porosity of the different freeze-dried HGGs

Once the crosslinking degree of the different HGGs was analyzed,

their apparent porosity was calculated and their morphology was

studied by SEM Fig 3 shows the images obtained from all samples once freeze-dried and cross-sectioned, while Table 1 shows HGG mean apparent porosity and the average diameter of the hydrogel macro- and

micropores, determined via image analysis

As can be noticed in both, Fig 3 and Table 1, the macro- and mi-cropores of the HGGAB samples were bigger than those of the HGGPBS

samples, which were less porous In addition, as can be observed in the images, HGGs prepared in PBS had more micropores than those syn-thesized in AB, which again revealed their greater degree of crosslinking

Likewise, regarding the diameter of the macro- and micropores of the HGGs prepared in AB with different concentrations of L-Cys, it should be noted that differences were not statistically significant in the case of macropores, but they were in the case of micropores, since those of the HGGsAB[1.5LCys] were smaller than the micropores of the other

hydrogels according to the post hoc analysis (Tukey test) that was later performed (p < 0.05) On the contrary, the differences in the size of the

macropores of the HGGsPBS were more remarkable than those of the micropores In this manner, the diameter of the micropores of all HGGsPBS was very similar, although as the concentration of L-Cys used in

Table 1

HGG apparent porosity (%) and mean diameter (μm) ± SD of the macro- and

micropores of the different hydrogel samples, once freeze-dried, determined via

SEM image analysis

Sample Porosity (%) Mean macropore size Mean micropore size

HGG AB [1.5LCys] 97.93 ± 1.4 553.1 ± 186.6 μm 325.9 ± 95.5 μm

HGG AB [3LCys] 96.53 ± 2.1 550.3 ± 155.5 μm 215.7 ± 69.0 μm

HGG AB [4.5LCys] 95.88 ± 1.7 561.0 ± 193.7 μm 225.6 ± 32.8 μm

HGG PBS [1.5LCys] 93.01 ± 0.9 442.3 ± 95.9 μm 123.2 ± 48.4 μm

HGG PBS [3LCys] 91.80 ± 1.6 354.3 ± 161.0 μm 123.0 ± 74.7 μm

HGG PBS [4.5LCys] 90.50 ± 1.8 390.0 ± 91.9 μm 127.2 ± 48.0 μm

Fig 4 (A) IR spectra of GG and the different HGGs in the 900–1800 cm− 1 (left) and 1800–4000 cm− 1 (right) ranges; (B) TG curves obtained for GG and the

different HGGs

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hydrogel synthesis increased, they had greater number of micropores

3.6 Chemical structure of the different HGGs

As can be seen in Fig 4(A), all GG characteristics bands within the

900–4000 cm− 1 range could be distinguished in the spectra of the

different HGGs In this manner, GG-specific peaks were observed at

1032 cm− 1 (–C–O–C– stretching), 1600 cm− 1 (C––O stretching

vi-brations), 2920 cm− 1 (–CH stretching) and 3400 cm− 1 (–OH

stretch-ing) in all samples (Lee et al., 2020) There were no significant

differences between the spectra of the HGGsAB and those of the HGGsPBS

Nevertheless, when comparing GG spectrum to the spectra of the

hydrogels, some alterations (marked in red in Fig 4[A]) could be

appreciated, possibly indicative of HGG successfully crosslinking with L-

Cys via EDC/NHS reaction Herein, HGGs had a peak at 1560–1562 cm− 1

that may correspond to the –CONH– amide bond formation between

GG –COOH and L-Cys –NH groups, and which was not present in GG

spectrum (Panczyszyn et al., 2021) The band at 1375 cm− 1, which

could correspond to the C–H bending and which was marked in the GG

spectrum (Criado et al., 2016), disappeared in the spectra of all HGGs

Finally, the characteristic peak of the -SH group was detected at 2530

cm− 1, and the peaks related to -CH2 vibrations at 2920–2929 cm− 1 were

more pronounced for the HGGs in comparison with GG, which may

confirm the thiolation of the hydrogels after L-Cys crosslinking (George

et al., 2020; Xu et al., 2021)

3.7 Thermal stability of the different HGGs

A TGA of the six different types of HGGs prepared was performed to

evaluate their thermal stability and mass loss and, thus, further

corroborate their crosslinking degree, since differences in degradation

temperatures can give some provision about polymer crosslinking TG

curves obtained for them can be seen in Fig 4(B), along with the GG

curve

As can be noticed in Fig 4(B), both GG and all HGGs showed a two- step thermogram, where the first stage of minor weight loss occurred in the 50–100 ◦C range This weight loss was likely caused by the evapo-ration of the adsorbed buffer/H2O in the samples Thus, it may be directly related to HGG swelling capacity (Ding et al., 2021; Karthika & Vishalakshi, 2015) and, for this reason, it was greater for the HGGAB

samples (11.1–14–4 %) than for the HGGPBS samples (8.5–11.0 %) and

GG (8.6 %) Likewise, HGGs crosslinked with lower L-Cys concentrations lost greater weight than those prepared with higher concentrations of the crosslinker, fact that showed again that L-Cys concentration in samples had an inverse relationship with the swelling capacity of the hydrogels and, consequently, with their porosity

On the other hand, the second stage of weight loss, which occurred in the 250–300 ◦C range, could account for GG degradation and the sub-sequent destruction of the whole hydrogel network structure (Ding

et al., 2021; Karthika & Vishalakshi, 2015) At this stage, HGGAB [1.5L-Cys], HGGAB[3LCys] and HGGAB[4.5LCys] samples lost about 50.6 %,

53 % and 53.7 % of weight, while HGGPBS[1.5LCys], HGGPBS[3LCys] and HGGPBS[4.5LCys] samples lost about 30.8 %, 32.4 % and 33.3 % of weight, respectively Thereby, the overall trend showed that the greater the degree of HGG crosslinking, the smaller their rate of weight loss and the better their thermal stability

3.8 Compression modulus of HGGs[3LCys]

The swelling and deswelling capacity of the hydrogels, which is determined by their crosslinking degree, governs drug release In this way, greater crosslinking degrees reduce hydrogel pore size and desw-elling capacity and decrease the overall diffusion of the drugs through the polymer networks (Khan & Ranjha, 2014; Sivakumaran et al., 2013) Therefore, based on the results obtained up to this point, it was considered that using HGGs[1.5LCys] could lead to a quick burst release

Fig 5 Degradation rate of HGGAB[3LCys] and HGGPBS[3LCys] samples after incubation with lysozyme solutions (1 mg/ml) at 37 ◦C for 9 days

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of PTX due to their larger pore size (Sivakumaran et al., 2013), while

PTX release from HGGs[4.5LCys] may be too slow because of their

elevated number of micropores Herein, those HGGs crosslinked with 3

mg/ml L-Cys were regarded to be the most suitable hydrogels to achieve

proper, local PTX release, and they were chosen to perform subsequent

assays

Therein, mechanical properties of the HGGs[3LCys] were analyzed

using static compression measurements The average Young's modulus

of both HGGsAB[3LCys] and HGGsPBS[3LCys] was found to be 86.5 ±

12.9 KPa and 95.9 ± 7.8 KPa, respectively Despite being close values (p

> 0.05), slightly increased mechanical strength in HGGsPBS was

ex-pected because of their higher degree of crosslinking In any case, the

compression elastic moduli of both hydrogels were in the range of the

modulus compression elasticity of most biological tissues that are soft

viscoelastic materials (0.1–100 KPa) (Shpaisman et al., 2012), so they

could meet the requirements to potentially be applied in vivo in the

future

3.9 Enzymatic degradation rate of HGGs[3LCys]

Before proceeding to load HGGs[3LCys] with the PTX:βCD com-plexes, their biosuitability was first analyzed using enzymatic degrada-tion assays The results obtained when investigating the degradadegrada-tion behavior of the HGGsAB[3LCys] and the HGGsPBS[3LCys] after incuba-tion with lysozyme soluincuba-tions can be seen in Fig 5

As can be observed in Fig 5, the weight of both hydrogel types decreased gradually with incubation time increasing, which proved their biodegradability Nonetheless, compared to HGGsPBS[3LCys],

Fig 6 (A) Results of the MTT assays performed with HS5 and BT474 cells to assess HGG biocompatibility Cells were exposed to both HGGsAB[3LCys] and HGGsPBS[3LCys] (23.1 % [v/v]), and their relative viability was compared with that of an untreated control The results shown are the average viability values ± SD

of three independent samples; (B) CLSM images of HS5 and BT474 cells 24 h after exposure to HGGs[3LCys] (23.1 % [v/v]) Cell survival and death were assessed by

using calcein AM (green) and propidium iodide (red)

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