1. Trang chủ
  2. » Tất cả

Effects of acitretin on semen quality and reproductive hormone levels in patients with psoriasis vulgaris

4 1 0
Tài liệu đã được kiểm tra trùng lặp

Đang tải... (xem toàn văn)

THÔNG TIN TÀI LIỆU

Thông tin cơ bản

Tiêu đề Effects of Acitretin on Semen Quality and Reproductive Hormone Levels in Patients with Psoriasis Vulgaris
Tác giả Hui Liu, Jie Li, Li Yu
Trường học Department of Dermatology, 210 Hospital of People's Liberation Army, Dalian
Chuyên ngành Dermatology
Thể loại Original Article
Năm xuất bản 2016
Thành phố Dalian
Định dạng
Số trang 4
Dung lượng 185,9 KB

Các công cụ chuyển đổi và chỉnh sửa cho tài liệu này

Nội dung

Effects of acitretin on semen quality and reproductive hormone levels in patients with psoriasis vulgaris lable at ScienceDirect DERMATOLOGICA SINICA xxx (2016) 1e4 Contents lists avai Dermatologica S[.]

Trang 1

ORIGINAL ARTICLE

Effects of acitretin on semen quality and reproductive hormone levels

in patients with psoriasis vulgaris

Hui Liu*, Jie Li, Li Yu

Department of Dermatology, 210 Hospital of People's Liberation Army, Dalian 116021, Liaoning, China

a r t i c l e i n f o

Article history:

Received: Aug 5, 2016

Revised: Sep 21, 2016

Accepted: Oct 31, 2016

Keywords:

acitretin

erythrocyte

innate immune

psoriasis

semen quality

sperm

a b s t r a c t Background: The aim of this study was to investigate the effect of different doses of acitretin (ACI) on the main parameters of semen, sperm morphology, and reproductive hormone levels in patients with psoriasis vulgaris

Methods: The main parameters of semen and the sperm morphology of 31 psoriatic patients were analyzed before and after treatment with different doses of ACI, and the changes in reproductive hormone levels were measured and compared with those of 14 healthy control individuals

Results: At treatment doses of 20 mg/d (n¼ 15) and 30 mg/d (n ¼ 16) of ACI, the semen volume, sperm motility, viability, concentration, and normal appearance percentage, and the serum reproductive hor-mone levels in the psoriatic patients showed no significant difference from those in healthy control individuals (n¼ 14) after 1 month and 3 months of treatment, as well as after withdrawing the treatment for 3 months (p> 0.05)

Conclusion: Different doses of ACI did not significantly affect semen quality in psoriatic patients at different treatment stages

Copyright© 2016, Taiwanese Dermatological Association Published by Elsevier Taiwan LLC This is an open access article under the CC BY-NC-ND license (http://

creativecommons.org/licenses/by-nc-nd/4.0/)

Introduction

Retinoids are a group of compounds that have similar structure and

biological activity as vitamin A Their bioactive isomers include

all-trans retinoid and 9-cis retinoid, which regulate cell proliferation

and differentiation, thus exhibiting bioactivities such as

keratosis, proliferation inhibition, antisebum synthesis,

anti-inflammation, and immunomodulation.1 Their molecular

struc-ture is composed of one cyclic group, one polyene side chain, and

one terminal polar group The change in any of the three parts could

result in compounds with different bioactivities Among the three

generations of retinoids,2acitretin (ACI) is the second-generation

derivative, and is suitable for treatment of all types of psoriasis

and keratosis Previously, the treatments of psoriasis with retinoids

were mostly focused on severe erythrodermic psoriasis and

pus-tular psoriasis,3,4which are types of psoriasis with significant side

effects Their indications kept on expanding, and low dose long-term treatment protocols were more widely used for psoriasis vulgaris, with satisfactory results and minimal side effects.5 Currently, it has been established that the major side effects of retinoids mainly manifest in muscle and bone, the central nervous system, liver, blood lipids, skin, and mucous membranes They have particularly potent teratogenic effects in women and female ani-mals, and women of childbearing age must use contraception during retinoid use and for at least 2 years after the withdrawal of retinoids.6 The specific effects of retinoids occur mainly in the cardiovascular system, central nervous system, and auditory sys-tem, and the occurrence rate could be as high as approximately 30% The embryotoxicity manifests as a high proportion of still-births and miscarriages, and the teratogenicity is still high a long time after withdrawal.6Thus, a consensus has been formed by to-day’s medical community that women of childbearing age must strictly use contraception during retinoid use The combination of sperm and egg, formation of a fertilized egg, and fetal development are all carried out in utero, so the effect of maternal factors on fetal development is self-evident Sperm cells are one of the most active and damage-susceptible cells in humans The father’s sperm qual-ity, especially the sperm concentration, mobilqual-ity, and morphology, affects the formation and development of the fertilized egg and the

Conflicts of interest: The authors declare that they have no financial or

non-financial conflicts of interest related to the subject matter or materials

discussed in this article.

* Corresponding author Department of Dermatology, 210 Hospital of People's

Liberation Army, Dalian 116021, Liaoning, China.

E-mail address: huiliudoc@126.com (H Liu).

Contents lists available atScienceDirect Dermatologica Sinica

j o u r n a l h o m e p a g e : h t t p : / / w w w d e r m - s i n i c a c o m

http://dx.doi.org/10.1016/j.dsi.2016.10.005

1027-8117/Copyright © 2016, Taiwanese Dermatological Association Published by Elsevier Taiwan LLC This is an open access article under the CC BY-NC-ND license ( http:// creativecommons.org/licenses/by-nc-nd/4.0/ ).

DERMATOLOGICA SINICA xxx (2016) 1e4

Trang 2

fetus It was reported that infertility caused by male factors

accounted for about half of the total number of cases of infertility.7

Thus, we must determine whether the increasingly widespread

usage of retinoids influences the reproductive system of male

pa-tients This has not received enough attention for a long time, and

the effect of retinoids on the semen quality and genital safety in

male psoriatic patients of childbearing age in China and abroad is

rarely reported Reproductive safety-targeted research wasfirstly

done by Parsch et al8in 1990, who used a manual method to detect

the semen quality changes in five patients after one retinoid

treatment (50 mg/d, 3e4 months) and concluded that retinoids had

no impact on sperm quality However, it is concerning that the

majority of the literature on the effects of retinoids on male genital

safety thereafter cited this report In recent years, foreign scholars

have begun to focus on the issue of retinoids’ application toward

male genital safety Because of ethical reasons, direct intervention

studies of ACI on the human male reproductive system in psoriatic

patients are very difficult Choosing animal models to obtain

tera-togenicity data and referring to the results of oral administration of

vitamin A and its major metabolites in the volunteers are the new

methods being used by certain research institutes to solve this

problem; however, the conclusions were different.9,10Using in vitro

tests, Pilkington and Brogden11found that high concentrations of

ACI could inhibit the fructose decomposition inside sperm,

signif-icantly decreasing the contents of lactic acid and CO2 Sigg et al12

conducted a small study on the effect of ACI on eight male

pa-tients, and reported no change in sperm concentration Although it

is still unknown whether ACI has residual effects in the semen of

male patients taking ACI during or after the treatment, it would not

constitute a risk to the fetus The maximum concentration of ACI

observed in human semen after taking ACI or etretinate was

12.5 ng/mL, namely about 125 ng/10 mL in semen However, there

is a warning that women of childbearing age and their spouses

must use contraception for at least 3 months before or after taking

isotretinoin in its package inserts Therefore, the aim of this study

was to investigate the changes of semen quality in 31 psoriatic

patients of childbearing age before and after treatment with

different doses of ACI Parameters analyzed included sperm

con-centration, motility, viability, and morphology, as well as levels of

three serum reproductive hormones: luteinizing hormone (LH),

follicle stimulating hormone (FSH), and serum testosterone (T) The

aim was to preliminarily explore the reproductive safety issues in

male patients using this kind of drug In this study, the detection of

physical indicators and parameters was done using a manual

method, and the sperm concentration and motility were analyzed

using a computerized automatic semen analysis system

Methods

General information

Thirty-one psoriatic patients were recruited from the in-patient

department of our hospital All were: young soldiers, aged 18e24

years, without any history of contact with radar or nuclear

radia-tion; unmarried with no children; with their primary disease as

psoriasis without orchitis, prostatitis, or varicocele; without

infec-tious diseases (such as hepatitis B, hepatitis C, or human

immu-nodeficiency virus); without endocrine or genetic diseases;

without familial high incidence of cancer; with a disease duration

of 3e48 months; and with a Psoriasis Area and Severity Index

(PASI) score of over eight points All of the patients were in the

progressive or stationary phase when semen examination was

performed The 31 patients were randomly divided into two groups

according to the doses of ACI The major parameters of semen

[sperm morphology and serum reproductive hormone levels (T,

FSH, and LH)] were observed before the treatment, 1 month after the treatment, 3 months after the treatment, and 3 months after withdrawal Group A: 15 cases, orally administered 20 mg once a day; Group B: 16 cases, orally administered 30 mg once a day The PASI score, age, disease duration, and body weight of the two groups showed no statistically significant difference (p > 0.05) All

of the patients were prohibited from undergoing any treatment during the withdrawal, except for external ointment therapy such

as tazarotene or mometasone The 14 healthy volunteers in the control group were young soldiers recruited from the surrounding area All were unmarried, without children, healthy, and aged 20e25 years All of the patients and volunteers were familiar with the study contents and gave informed consent

Semen tests All participants were asked to abstain from sexual activity for 2e7 days before the semen collection During the sampling, benzalko-nium bromide solution and saline were used to swab the urethra and surrounding skin Then, the semen was obtained in a sterile wide-mouth glass bottle by masturbation so as to avoid contami-nation and leakage The semen was liquefied, and the sperm count, motility, and morphological analyses were measured according to the standards described in the Laboratory Manual of WHO human semen and spermdcervical mucus interactions (fifth edition).13 The semen volume was measured, and the color, sticky degree,

or liquefaction were observed The pH was determined using pH test strips after the semen was liquefied A sperm quality automatic analyzer (SCA 5.2; Microptic, Barcelona, Spain) was used to mea-sure the sperm concentration, viability, and motility.14

Sperm morphology observations The results of sperm morphology observations were manually counted using the Diff-Quik method.15

Detection of sperm nuclear DNA fragments Sperm nuclear DNA fragments were detected usingflow cytometry (Accuri C6; Becton Dickinson Immunocytometry Systems, San Jose,

CA, USA).16 Detection of reproductive hormones by radioimmunoassay The levels of T, FSH, and LH were detected using the BHP9507 chemiluminescence immunoassay analyzer and kit (Yamasa Shoyu Co., Ltd., Chiba, Japan),17with the intrabatch error< 5%

Statistical analysis The comparison between different ACI dose groups, different treatment period groups, and the control group was performed using the t test The measurement data were expressed as x±SD, and all the data were processed using SPSS13.0 statistical package (SPSS Company, Chicago, Illinois, USA), witha¼ 0.05 set as the test level

Ethical approval This study was conducted in accordance with the declaration of Helsinki This study was conducted with approval from the Ethics Committee of 210 Hospital of People's Liberation Army Written informed consent was obtained from all participants

H Liu et al / Dermatologica Sinica xxx (2016) 1e4

Trang 3

Comparison of routine semen tests

The concentration, motility, viability, and morphology of sperm, the

semen volume, and the pH in Groups A and B showed no significant

difference compared with those in the control group before the

treatment or 1 month and 3 months after the treatment (p> 0.05,

Table 1)

Comparison of sperm nuclear DNA fragments

The redfluorescence intensity (FL1-H) and red fluorescence

in-tensity/totalfluorescence intensity (at) in Groups A and B showed

no significant difference compared with those in the control group

before the treatment or 1 month and 3 months after the treatment

(p> 0.05,Table 2)

Comparison of reproductive hormone levels

The levels of T, FSH, and LH in Groups A and B showed no significant

difference compared with those in the control group before the

treatment or 1 month and 3 months after the treatment (all

p> 0.05,Table 3)

Comparison of semen quality and reproductive hormone levels

The semen quality indexes and levels of T, FSH, and LH in Groups A

and B showed no significant difference compared with those in the

control Group 3 months after withdrawal (p> 0.05,Table 4)

Discussion

It has been confirmed that ACI has strong teratogenic effects in

humans and animals Thus, women of childbearing age must use

contraception for at least 2 years after the withdrawal of

treat-ment.6There is a large number of studies on the teratogenicity and

embryotoxicity of retinoids after their oral and topical

adminis-tration in animals, but retinoid risk assessments in humans mainly

include indirect data of pharmacokinetic analysis in animals and

humans after systematic administration experiments and topical

retinoid application in humans, which also referred to the results of

in vitro teratogenic tests in animals and humans.18e20 The most

direct determinants of teratogenic effects depend on the

concentration-time relationship of drugs in target tissues, the

drugs’ physicochemical and structure-activity properties, and their

concentration and affinity in the terminal target organs.21Although

pharmacological and pharmacokinetic preventive measures have

been proposed, they are still widely used clinically because of their

efficacy.22Currently, there are few studies on the effects of retinoids

on the male reproductive system These studies are based on tests and literature, and most of them consider retinoid treatment to be safe from the viewpoint of andrology Given the wide clinical usage

of retinoids, clarification of whether retinoids affect the reproduc-tive system in male patients of childbearing age is urgently needed

Table 1 Comparison of semen quality between psoriatic patients and normal controls in different periods ðx±sÞ.

Group Course (mo) Semen volume

(mL)

pH Total motility

(PRþNP) (%)

Ratio of live sperm (%)

Sperm concentration (10 6 /mL)

Percentage of sperm with normal morphology (%) A

(20 mg/d, n ¼ 15)

0 3.4 ± 0.4 7.5 ± 0.1 63.1 ± 3.8 79.8 ± 13.6 46.9 ± 10.2 3.9 ± 0.7

1 3.3 ± 0.5 7.4 ± 0.3 64.1 ± 4.0 81.1 ± 14.1 48.1 ± 11.0 3.8 ± 0.3

3 3.4 ± 0.2 7.5 ± 0.1 63.9 ± 2.7 79.6 ± 12.7 47.5 ± 9.8 3.8 ± 0.2 B

(30 mg/d, n ¼ 16)

0 3.3 ± 0.6 7.4 ± 0.2 64.0 ± 2.9 80.1 ± 11.7 48.0 ± 9.9 3.8 ± 0.4

1 3.4 ± 0.1 7.6 ± 0.1 63.7 ± 3.1 81.4 ± 12.9 46.9 ± 8.9 3.7 ± 0.6

3 3.4 ± 0.2 7.5 ± 0.3 63.1 ± 4.1 79.2 ± 14.0 47.1 ± 11.3 3.9 ± 0.5 Control

(n ¼ 14)

0 3.4 ± 0.3 7.5 ± 0.2 63.8 ± 3.6 80.4 ± 12.2 47.6 ± 10.9 3.8 ± 0.5

NP ¼ nonprogressive sperm; PR ¼ progressive sperm.

Table 2 The variety of DNA fragments determination of sperm nuclear for men between psoriatic patients and normal controls in different periods ðx±sÞ Group Course (mo) Red fluorescence

intensity (FL1-H)

Red fluorescence intensity/total fluorescence intensity (at )

A (20 mg/d, n ¼ 15) 0 13.92 ± 1.03 9.92 ± 3.05

1 13.77 ± 0.90 10.26 ± 3.90

3 14.18 ± 1.16 9.90 ± 2.37

B (30 mg/d, n ¼ 16) 0 14.47 ± 1.40 10.77 ± 4.01

1 13.92 ± 1.07 9.89 ± 2.64

3 13.82 ± 1.01 9.90 ± 6.31 Control (n ¼ 14) 0 14.13 ± 1.48 10.10 ± 5.61

Table 3 The variety of the genital hormone for men between psoriatic patients and normal controls in different treatment stage ðx±sÞ.

Group Course (mo) T (ng/mL) FSH (ng/mL) LH (ng/mL)

A (20 mg/d, n ¼ 15) 0 744 ± 189 7.2 ± 1.4 2.7 ± 1.1

1 746 ± 240 7.3 ± 1.5 2.8 ± 1.7

3 739 ± 171 7.3 ± 0.9 2.7 ± 1.6

B (30 mg/d, n ¼ 16) 0 751 ± 201 7.3 ± 1.0 2.8 ± 1.3

1 737 ± 223 7.3 ± 0.9 2.7 ± 1.4

3 742 ± 193 7.4 ± 1.3 2.9 ± 1.9 Control (n ¼ 14) 0 740 ± 235 7.3 ± 1.6 2.8 ± 1.8 FSH ¼ follicle stimulating hormone; LH ¼ luteinizing hormone; T ¼ testosterone.

Table 4 Comparison of semen quality and genital hormone between psoriatic pa-tients and normal controls after 3 months of withdrawal ðx±sÞ Index A (20 mg/d,

n ¼ 14)

B (30 mg/d,

n ¼ 13)

Control (n ¼ 14) Semen volume (mL) 3.3 ± 0.4 3.4 ± 0.2 3.4 ± 0.3

pH 7.4 ± 0.2 7.3 ± 0.3 7.5 ± 0.2 Total mobility (PR þ NP) (%) 63.7 ± 2.7 62.8 ± 3.1 63.8 ± 3.6 Ratio of live sperm (%) 81.1 ± 11.3 80.9 ± 12.9 80.4 ± 12.2 Sperm concentration ( 10 6 /L) 48.6 ± 9.9 47.1 ± 10.2 47.6 ± 10.9 Ratio of sperm with normal

morphology (%)

3.8 ± 0.3 3.7 ± 0.6 3.8 ± 0.5

T (ng/mL) 737 ± 179 743 ± 201 740 ± 235 FSH (ng/mL) 7.3 ± 1.3 7.4 ± 0.6 7.3 ± 1.6

LH (ng/mL) 2.7 ± 1.6 2.8 ± 1.3 2.8 ± 1.8 Red fluorescence intensity 14.60 ± 1.45 13.93 ± 1.36 14.13 ± 1.48 Red fluorescence intensity/total

fluorescence intensity

9.94 ± 5.61 10.30 ± 3.44 10.10 ± 5.61

FSH ¼ follicle stimulating hormone; LH ¼ luteinizing hormone; NP ¼ nonprog-ressive sperm; PR ¼ progressive sperm; T ¼ testosterone.

H Liu et al / Dermatologica Sinica xxx (2016) 1e4

Trang 4

As one of the most active and damage-susceptible cells in vivo,

the development and maturation of sperm cells are complex These

processes mainly occur in testes and seminiferous tubes, and

consist of three phases that together last 2e3 months: the

prolif-eration and division of spermatogonia, reduction division of

sper-matocytes, and spermatogenesis.7 Routine tests and analysis of

semen and sperm are a direct and objective experimental method

in clinical practice The comprehensive assessment of semen

vol-ume and liquefaction time, as well as sperm concentration, motility,

viability, morphology, and other indexes could be used as an

important basis for understanding male fertility The male

repro-ductive hormone levels depend on the

hypothalamic-pituitary-gonadal axis, and changes to this axis are directly related to

changes in sperm concentration, motility, and morphology seen in

semen analysis

To understand the effects of different doses of ACI on sperm, we

compared the semen volume, motility, survival ratio, sperm

con-centration, percentage of sperm with normal morphology,

damaged DNA fragment index, and certain genital hormones

among 15 psoriatic patients administered 20 mg of ACI/d, 16

pso-riatic patients administered 30 mg of ACI/d, and 14 healthy

vol-unteers before the treatment and 1 month and 3 months after

treatment The results showed that ACI can improve the clinical

symptoms of psoriatic patients, but different doses of ACI did not

cause significant changes in sperm concentration, sperm

morphology, total sperm count, sperm motility, or reproductive

hormone levels 1 month or 3 months after treatment Three

months after withdrawal of treatment (one period of

spermato-genesis), we rechecked the semen quality and reproductive

hor-mone levels of 27 patients (3 patients retired from active military

service, and 1 patient quit) The results did not reveal any

abnor-malities, indicating that long-term oral treatment with ACI did not

affect spermatogenesis and that ACI did not affect sperm

concen-tration, reduce sperm motility, or affect sperm morphology ACI had

no significant effect on the male hypothalamic-pituitary-gonadal

axis or its regulated reproductive hormone levels during or after

treatment Compared with the previous experiments,8the patients

in this study were younger and the volunteers were young soldiers

without psoriasis; therefore, the study samples were easily

managed and followed up, and their compliance was better The

sample size was larger than in previous studies, the treatment

protocol used conventional doses (no more than 30 mg/d), and the

conclusions obtained were consistent Therefore, the study

confirmed that long-term treatment with therapeutic doses of

retinoids has no adverse effects on reproductive function in

pso-riatic male patients

ACI has confirmed efficacy in the treatment of psoriasis, and

despite its relatively many side effects, it shows no adverse effects

in the reproductive system of male patients of childbearing age

during treatment Many issues regarding the use of ACI have not

been completely understood, such as post-treatment long-term

follow-up and reproductive safety issues of other drugs used

concomitantly with ACI (such as isotretinoin) in male patients of

childbearing age Confirming whether ACI affects the reproductive

system of male psoriatic patients of childbearing age is an impor-tant public health problem that requires urgent solutions Answering this question would provide a strong theoretical basis for clinical and scientific application and safety assessment of retinoids

References

1 Alsenaid A, Eder I, Ruzicka T, Braun-Falco M, Wolf R Successful treatment of nail lichen planus with alitretinoin: report of 2 cases and review of the liter-ature Dermatology 2014;229:293e6

2 Kong R, Cui Y, Fisher GJ, et al A comparative study of the effects of retinol and retinoic acid on histological, molecular, and clinical properties of human skin.

J Cosmet Dermatol 2016;15:49e57

3 Tang MM, Spanou Z, Tang H, Schibler F, Pelivani N, Yawalkar N Rapid down-regulation of innate immune cells, interleukin-12 and interleukin-23 in generalized pustular psoriasis with infliximab in combination with acitretin Dermatology 2012;225:338e43

4 Gallo E, Llamas-Velasco M, Dauden E, García-Diez A Refractory generalized pustular psoriasis responsive to a combination of adalimumab and acitretin Int

J Dermatol 2013;52:1610e1

5 Popadic S, Nikolic M Pustular psoriasis in childhood and adolescence: a 20-year single-center experience Pediatr Dermatol 2014;31:575e9

6 Raguideau F, Mezzarobba M, Zureik M, Weill A, Ricordeau P, Alla F Compliance with pregnancy prevention plan recommendations in 8672 French women of childbearing potential exposed to acitretin Pharmacoepidemiol Drug Saf 2015;24:526e33

7 Rapino C, Battista N, Bari M, Maccarrone M Endocannabinoids as biomarkers of human reproduction Hum Reprod Update 2014;20:501e16

8 Parsch EM, Ruzicka T, Przybilla B, Schill WB Andrological investigations in men treated with acitretin Andrologia 1990;22:479e82

9 Cathryn AH, Michael DG Retinoic acids regulation of male meiosis Androgens 2013;20:217e23

10 Josephine B, Peter K Retinoic acid, meiosis and germ cell fate in mammals Development 2007;134:3401e11

11 Pilkington T, Brogden RN Acitretinda review of its pharmacology and thera-peutic use Drugs 2002;43:597e627

12 Sigg C, Bruckner-Tuderman L, Gilardi S Andrological investigations in patients treated with etretin Dermatologica 1987;175:48e9

13 World Health Organization WHO laboratory manual for the Examination and processing of human semen 5th ed Geneva: WHO; 2010 p 1e286

14 Dorado J, Rijsselaere T, Mu ~noz-Serrano A, Hidalgo M Influence of sampling factors on canine sperm motility parameters measured by the Sperm Class Analyzer Syst Biol Reprod Med 2011;57:319e24

15 Tavares RS, Silva AF, Lourenço B, Almeida-Santos T, Sousa AP, Ramalho-Santos J Evaluation of human sperm chromatin status after selection using a modified Diff-Quik stain indicates embryo quality and pregnancy outcomes following in vitro fertilization Andrology 2013;1:830e7

16 Novotny J, Aziz N, Rybar R, et al Relationship between reactive oxygen species production in human semen and sperm DNA damage assessed by Sperm Chromatin Structure Assay Biomed Pap Med Fac Univ Palacky Olomouc Czech Repub 2013;157:383e6

17 Safarinejad MR Clinical significance of reproductive hormone test J Urol 2009;182:237e48

18 Nya-Ngatchou JJ, Arnold SL, Walsh TJ, et al Intratesticular 13-cis retinoic acid is lower in men with abnormal semen analyses: a pilot study Andrology 2013;1: 325e31

19 Hogarth CA, Arnold S, Kent T, Mitchell D, Isoherranen N, Griswold MD Proc-essive pulses of retinoic acid propel asynchronous and continuous murine sperm production Biol Reprod 2015;92:1e11

20 Busada JT, Chappell VA, Niedenberger BA, et al Retinoic acid regulates Kit translation during spermatogonial differentiation in the mouse Dev Biol 2015;397:140e9

21 Zheng L, Xu T, Li D, Zhou J A representative retinoid X receptor antagonist UVI3003 induced teratogenesis in zebrafish embryos J Appl Toxicol 2015;35: 280e6

22 Monfrecola G, Baldo A Retinoids and phototherapy for psoriasis J Rheumatol Suppl 2009;83:71e2

H Liu et al / Dermatologica Sinica xxx (2016) 1e4

Ngày đăng: 24/11/2022, 17:54

TÀI LIỆU CÙNG NGƯỜI DÙNG

TÀI LIỆU LIÊN QUAN

🧩 Sản phẩm bạn có thể quan tâm