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states Arizona, California, Colorado, Idaho, Montana, Nev-ada, New Mexico, Texas, Utah, Oregon, Washington, and Wyoming; Figure 2 Flather et al., 1994.. states Hawaii is the hot spot of

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states (Arizona, California, Colorado, Idaho, Montana,

Nev-ada, New Mexico, Texas, Utah, Oregon, Washington, and

Wyoming; Figure 2) (Flather et al., 1994) However, among

U.S states Hawaii is the hot spot of endangerment: about 950

native plant species of Hawaii are extinct, endangered, or

threatened (Raven et al., 1986)

Endangerment patterns vary by habitat type.Flather et al.

plants in the continental United States are found in

range-lands Twenty-five percent of endangered plant species were

associated with barren land or rocky habitats However, wet-lands, which comprise only 5% of the land base, contain 15%

of the listed species

Endangerment patterns also vary by region and land-use characteristics In northern California, southern Nevada/ Sonoran Basin, and the eastern Gulf Coast, 450% of the listed species are associated with wetland habitats U.S Department

of Defense lands, which comprise only 3.4% of federally ad-ministered lands, contain 26% of the listed species The high number of rare plants in the U.S Southwest when combined

Canada 46

Greenland 3

United Kingdom 24 Ireland 3 France 78 Portugal 77 Morocco 107

Mauritania 13 Senegal 71 Liberia 96 Ghana 205 Cameroon 529 Gabon 219

Nigeria 286

Niger 33 Mali 49

Algeria 73

Spain 135

Norway 12 Sweden 11 Finland 8

Italy 83 Turkey 54

Chad 34

Egypt 30 Sudan 80 DR Congo 217 Angola 128 Namibia 216 Botswana 58 South Africa 300

Mauritius 100 Madagascar 386 Tanzania 470Kenya 290 Somalia 105 Yemen 340 Oman 18

Saudi Arabia 18

Iran 22

Kazakhstan 55

India 424

China 686

Russian Federation 56

Mongolia 13

Thailand 258

Malaysia 1,234 Indonesia 754 Australia

249 New Caledonia346

New Zealand 60

Papua New

Taiwan 136 Philippines 373 Vietnam 303 Sri Lanka

334

Japan 88 United

413 Mexico

462 Cuba 193 Dominican Republic

47 Puerto Rico 69 Venezuela 208 Guyana 67 Suriname 55

Brazil 623

Jamaica 300 Guatemala 132

French

Polynesia

158

Costa Rica 194 Panama 329 Ecuador 2,549 Colombia 352

Peru 379 Bolivia 113 Chile

62 Argentina

94 Uruguay 8 Paraguay 38

Figure 1 Distribution of rare plants in the world Data from IUCN (2010) IUCN Red List of Threatened Species Version 2010.3 http://

www.iucnredlist.org Downloaded on 05 October 2010

WASHINGTON

OREGON

IDAHO MONTANA

WYOMING

NEVADA

CALIFORNIA

ARIZONA

UTAH

COLORADO

NEW MEXICO

TEXAS OKLAHOMA KANSAS NEBRASKA

SOUTH DAKOTA

NORTH DAKOTA MINNESOTA

IOWA WISCONSIN

ILLINOIS MISSOURI

ARKANSAS

LOUISIANA MISSISSIPPI ALABAMAGEORGIA

FLORIDA

SOUTH CAROLINA

NORTH CAROLINA TENNESSEE

KENTUCKY INDIANA OHIO

MICHIGAN VERMONT MAINE

NEW HAMPSHIRE MASSACHUSETTS RHODE ISLAND CONNECTICUT NEW JERSEY DELAWARE MARYLAND

None known 1−4 5−10 11−30

>30

VIRGINIA

WEST VIRGINIA PENNSYLVANIA NEW YORK

Figure 2 Distribution of rare plants in the continental United States The ‘‘top 10’’ continental states for rare plants include six western states, and California clearly dominates Reproduced from Flather CH, Knowles MS, and Kendall IA (1998) Threatened and endangered species

geography BioScience 48: 365–376.

Endangered Plants 207

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