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Tiêu đề Inhibition of cobalamin-dependent methionine synthase by substituted benzo-fused heterocycles
Tác giả Elizabeth C. Banks, Stephen W. Doughty, Steven M. Toms, Richard T. Wheelhouse, Anna Nicolaou
Trường học School of Pharmacy, University of Bradford
Chuyên ngành Pharmacy
Thể loại Research article
Năm xuất bản 2006
Thành phố Bradford
Định dạng
Số trang 13
Dung lượng 1,35 MB

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Nội dung

MetS is therefore intimately linked to important biochemical Keywords benzimidazole; benzothiadiazole; inhibition; methionine synthase; molecular modelling Correspondence A.. Kinetic ana

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by substituted benzo-fused heterocycles

Elizabeth C Banks1, Stephen W Doughty2,*, Steven M Toms1, Richard T Wheelhouse1

and Anna Nicolaou1

1 School of Pharmacy, University of Bradford, UK

2 School of Pharmacy, Centre for Biomolecular Sciences, University of Nottingham, UK

Methionine synthase (MetS)

(5-methyltetrahydrofolate-homocysteine transmethylase)

two established mammalian enzymes that utilize a

bio-logically active cobalamin derivative [methylcobalamin

(CH3-Cbl)] as a cofactor [1] MetS catalyses the transfer

of the methyl group from 5-methyltetrahydrofolate to

homocysteine via the CH3-Cbl cofactor, with cycling

of cobalamin between the +1 [Cbl(I)] and +3 [Cbl(III)]

valency states (Fig 1) Studies on the Escherichia coli

and Homo sapiens cobalamin-dependent MetS have

revealed that it is a large, conformationally flexible

pro-tein, consisting of four functional domains arranged

in a linear manner Each one of these domains binds a

different substrate or cofactor In detail, the N-terminal module is the homocysteine (Hcy)-binding domain; the second domain binds 5-methyltetrahydrofolate, the third domain binds CH3-Cbl; and the fourth domain (C-term-inal module) binds S-adenosyl-methionine (S-AdoMet),

an allosteric cofactor required for reductive reactivation [2] X-ray crystal structures of the cobalamin-, S-AdoMet-and 5-methyltetrahydrofolate-binding sites have only been reported for the bacterial enzyme [3–5]

The reaction products methionine and tetrahydro-folate are further metabolized through the one-carbon methionine transmethylation and folate cycles MetS

is therefore intimately linked to important biochemical

Keywords

benzimidazole; benzothiadiazole; inhibition;

methionine synthase; molecular modelling

Correspondence

A Nicolaou, School of Pharmacy, University

of Bradford, Richmond Road, Bradford BD7

1DP, UK

Fax: +44 1274 235600

Tel: +44 1274 234717

E-mail: a.nicolaou@bradford.ac.uk

*Present address

Faculty of Health and Biological Sciences,

School of Pharmacy, University of

Notting-ham Malaysia Campus, Jalan Broga, 43500

Semenyih, Selangor Darul Ehsan, Malaysia

(Received 14 July 2006, revised 7 November

2006, accepted 9 November 2006)

doi:10.1111/j.1742-4658.2006.05583.x

The cobalamin–dependent cytosolic enzyme, methionine synthase (EC.2.1.1.13), catalyzes the remethylation of homocysteine to methionine using 5-methyltetrahydrofolate as the methyl donor The products of this remethylation – methionine and tetrahydrofolate – participate in the active methionine and folate pathways Impaired methionine synthase activity has been implicated in the pathogenesis of anaemias, cancer and neurological disorders Although the need for potent and specific inhibitors of methion-ine synthase has been recognized, there is a lack of such agents In this study, we designed, synthesized and evaluated the inhibitory activity of a series of substituted benzimidazoles and small benzothiadiazoles Kinetic analysis revealed that the benzimidazoles act as competitive inhibitors of the rat liver methionine synthase, whilst the most active benzothiadiazole (IC50¼ 80 lm) exhibited characteristics of uncompetitive inhibition A model of the methyltetrahydrofolate-binding site of the rat liver methionine synthase was constructed; docking experiments were designed to elucidate,

in greater detail, the binding mode and reveal structural requirements for the design of inhibitors of methionine synthase Our results indicate that the potency of the tested compounds is related to a planar region of the inhibitor that can be positioned in the centre of the active site, the presence

of a nitro functional group and two or three probable hydrogen-bonding interactions

Abbreviations

CH 3 -Cbl, methylcobalamin; DHPS, dihydropteroate synthase; Hcy, homocysteine; IC 50 , half-inhibitory concentration; MeTr, methyltransferase protein; MetS, methionine synthase; S-AdoMet, S-adenosylmethionine.

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pathways These include the reactions of

trans-sulfura-tion through the productrans-sulfura-tion of homocysteine,

biologi-cal methylations of DNA, lipids and proteins, and

polyamine biosynthesis through the production of

methionine and S-AdoMet [6,7] Furthermore, MetS is

the only human enzyme that metabolizes

methyltetra-hydrofolate to tetramethyltetra-hydrofolate, thereby facilitating the

recycling of this major form of folates to other

bio-active folates that provide one-carbon units for purine

and pyrimidine synthesis Impaired function of MetS

has been linked to megaloblastic anaemias and

neuro-logical disorders [8], atherosclerosis [7] and

carcinogen-esis [9,10]

Although the need to develop inhibitors of MetS as

drug candidates has long been recognized [11], there

are a limited number of reports on agents inhibiting

this enzyme The anaesthetic gas N2O is possibly the

only selective inhibitor of MetS reported to date, its

action mediated through the oxidation of the

cobala-min cofactor [12] Other compounds that have been

shown to inhibit this enzyme are the cell-signalling

molecule nitric oxide [13,14], chloroform and carbon

tetrachloride [15], methylmercury [16], ethanol and

acetaldehyde [17], hydrazine [18], S-AdoMet

deriva-tives [19] and a series of cobalamin analogues [20]

Polyamines have been shown to stimulate MetS

activity [21], whilst methotrexate has been shown to

indirectly inhibit the enzyme in vivo through depletion

of its substrate, 5-methyltetrahydrofolate [22]

In a new strategy for discovering specific inhibitors

of MetS, drug-like, benzo-fused heterocycles that

mimic substructures of 5-methyltetrahydrofolate have

been evaluated in a cell-free system The inhibitory

activity and mechanism of action have been probed by

kinetic studies using purified rat liver enzyme, whilst a

diamines were prepared by selective reduction of nitroanilines [23] and cyclized with formic acid [24] to give the desired substituted benzimidazoles The benz-imidazoles 1a–b and the benzothiadiazoles 2a–c were commercially available

All the compounds were tested against highly puri-fied rat liver MetS [14] and the half-inhibitory con-centrations (IC50) are presented in Table 1 The benzimidazoles 1c, 1h and 1k, and the nitrobenzothiad-iazole 2b, gave IC50 values close to or below 100 lm, with 2b being the most potent inhibitor (IC50¼

80 lm) From these results it was apparent that the presence of a nitro group at the 5-position was associ-ated with stronger inhibition (1c compared with 1d; 1h with 1i) and this was positively associated with the presence of the 3-methoxy group (1c compared with 1h) However, the aminobenzimidazole 1k showed a marginally stronger inhibition than the corresponding nitrobenzimidazole 1j This result may indicate that there is more than one mode of interaction with the active site that affects the activity of more-highly sub-stituted molecules such as 1j and 1k Furthermore, the presence of an N-methyl group on the benzimidazole ring (position similar to the one in the substrate 5-methyltetrahydrofolate; Fig 1) was detrimental to the IC50 (comparing 1c with 1f and 1h with 1j) Finally, the inhibitory activity of the benzothiadiazoles was improved by the nitro substitution (2b compared with 2a or 2c)

To explore further the molecular mechanism of action of those two classes of substituted benzohetero-cycles, the kinetic parameters of inhibition were meas-ured Compounds 1c and 2b were chosen as being representative of each class as a result of their good inhibitory activity and availability Figure 2 shows the Lineweaver–Burk, Dixon and Cornish–Bowden plots for the uninhibited and inhibited reactions The Km

values for the uninhibited reaction were calculated to

be 25 lm for 5-methyltetrahydrofolate and 0.6 lm for homocysteine, both results being in fair agreement

Fig 1 The cobalamin-dependent methionine synthase catalysed

reaction Cbl(I), cob(I)alamin; CH 3 -Cbl, methylcobalamin; R,

ptero-glutamate.

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with previously published data for the pig liver enzyme

(16.8 and 2.16 lm, respectively) [25] The Lineweaver–

Burk plots for the inhibited reactions showed that the

nitrobenzimidazole 1c exhibits the characteristics of

mixed inhibition (Fig 2A) (Ki¼ 26 lm), whilst the

nitrobenzothiadiazole 2b is an uncompetitive inhibitor

of MetS with respect to 5-methyltetrahydrofolate

(Ki¼ 17 lm) (Fig 2B) These findings were confirmed

by the Dixon (Fig 2C,D) [26] and Cornish–Bowden

(Fig 2E,F) [27] plots for the two compounds When

1c and 2b were assessed with homocysteine as the

vari-able substrate, the Lineweaver–Burk, Dixon and

Corn-ish–Bowden plots indicated that both compounds

exhibited characteristics of mixed inhibition (Fig 3),

with the nitrobenzothiadiazole 2b presenting a strong

component of uncompetitive inhibition (Fig 3D) It

must also be noted that both compounds were very

weak inhibitors when assessed with respect to

homo-cysteine, with detectable inhibition noted mainly at

high concentrations of the inhibitors (0.5 and 1 mm;

Fig 3A)

The results of these studies suggest that the two

clas-ses of substituted benzo-fused heterocycles may act by

two distinct mechanisms The mixed inhibition

exhib-ited by the nitrobenzimidazole 1c is a pattern usually

observed in multisubstrate enzyme-catalysed reactions

such as MetS However, the uncompetitive inhibition,

shown by the nitrobenzothiadiazole 2b, indicates that

there is no reversible link between the inhibitor and

the variable substrates Plausible rationalization

includes the possibility that the relatively small nitro-benzothiadiazole may displace the dimethylbenzimidaz-ole side chain of the cobalamin-cofactor or that it may act on the binding site of the MetS allosteric cofactor, S-AdoMet, both effects which could explain the observed uncompetitive inhibition Furthermore, clo-sely related 1,2,3-benzothiadiazoles and 1,2,4-thiazoles act as potent electron acceptors in biological systems Thus, 1,2,4-thiadiazoles can be used to trap cysteine residues by mixed disulfide formulation [28] Alternat-ively, 1,2,3-benzothiazoles have been shown to inhibit cytochrome P450 metabolites by interference with elec-tron transport within the catalytic cycle of cytochrome P450 [29] Details of the potential binding and electron transfer events that may account for the uncompetitive inhibition of MetS by nitrobenzothiadiazole 2b are the subject of continuing investigation in this laboratory

To elucidate further the mechanism of action of the substituted benzo-fused heterocycles, to explore the interactions occurring at the binding site, and to develop a tool that could assist further optimization

of inhibitors, a molecular model of the methyltetra-hydrofolate-binding domain of the rat liver MetS was constructed In the absence of a high-resolution struc-ture of the methyltetrahydrofolate-binding site for the mammalian enzyme, a model based on the X-ray crys-tal structure of the methyltetrahydrofolate corrinoid iron-sulfur methyltransferase protein (MeTr) from Clostridium thermoaceticum, as determined by Doukov

et al [30], was constructed It has been suggested that

Table 1 Structures and half inhibitory concentrations (IC 50 ) of the series 1 and 2 substituted benzo-fused heterocycles IC 50 values were determined using highly purified rat liver methionine synthase.

N N

R X

Y

1 5

7

N S N

Z

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the methyltetrahydrofolate-binding domain of MetS,

and indeed of other methyltransferases, share

architec-tural similarities [30] We therefore constructed the rat

liver MetS methyltetrahydrofolate-binding site

(resi-dues 359–639) model based on the homology of this

protein with that of MeTr (residues 1–262) Using

information from the Brookhaven protein data bank, sequence homologues of the two proteins were obtained (Fig 4) This led to the deduction of a back-bone structure that was modelled to the conserved TIM barrel fold (triose phosphate isomerase type structure), a common feature for globular proteins that

D C

F E

Fig 2 Lineweaver–Burk plots for nitrobenzimidazole 1c (A) and nitrothiadiazole 2b (B), with respect to methyltetrahydrofolate (MTHF), at inhibitor concentrations of 1000 l M (·), 500 l M (m), 100 l M (d) and 0 l M (j) Dixon plots for nitrobenzimidazole 1c (C) and nitrothiadiazole 2b (D), and Cornish–Bowden plots for nitrobenzimidazole 1c (E) and nitrothiadiazole 2b (F), with respect to the inhibitor, at methyltetrahydro-folate (MTHF) concentrations of 11 l M (·), 22 l M (m), 67 l M (d) and 224 l M (j).

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has been predicted to occur in the pterin-binding site

of related methyltransferases [30] The nonconserved

sequences were altered and any insertions or deletions

were applied using the molecular modelling program,

sybyl

3 , to construct and refine the model A gradual refinement of the resulting structure was performed using minimization through application of the charmm

4 program and force field [31]

A

C

E

B

D

F

Fig 3 Lineweaver–Burk plots for nitrobenzimidazole 1c (A) and nitrothiadiazole 2b (B), with respect to homocysteine (Hcy), at inhibitor con-centrations of 1000 l M (·), 500 l M (m), 100 l M (d) and 0 l M (j) Dixon plots for nitrobenzimidazole 1c (C) and nitrothiadiazole 2b (D), and Cornish–Bowden plots for nitrobenzimidazole 1c (E) and nitrothiadiazole 2b (F), with respect to the inhibitor, at homocysteine (Hcy) concen-trations of 1.1 l M (·), 2.2 l M (m), 6.5 l M (d) and 11 l M (j).

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Docking of the substrate 5-methyltetrahydrofolate in

the model of the active site was based on the approach

followed by Doukov et al [30] to identify the

inter-actions between the pterin cofactor and MeTr This

approach was based on the assumption that the strong

structural homology of MeTr and the dihydropteroate

synthases (DHPS) allows the prediction of interactions

between the pterin ring and MeTr based on the

binding of hydroxymethylpterin pyrophosphate to

DHPS Following the same approach, and in order to

identify the orientation of the substrate when bound

to the active site, we superimposed the rat liver MetS

model on the experimentally determined structure of

hydroxymethlypterin pyrophosphate bound to DHPS

[32] The result of this approach indicated the

orienta-tion of 5-methyltetrahydrofolate in the MetS active

site Figure 5 shows the superimposed structures of

DHPS and 5-methyltetrahydrofolate The model of the

methyltetrahydrofolate-binding site of MetS with one

molecule of the substrate included was further

opti-mized using charmm, and the ligand was

parameter-ized using partial atomic charges and other parameters

obtained from quantum mechanic modelling

(Hartree-Fock 6–31G* within the Spartan PCPro package)

the ligand structure Electrostatic surfaces of the

meth-yltetrahydrofolate-binding site domain were generated

to show the size of the active site (Fig 6) The

negat-ively charged areas may indicate the need of the

inhib-itor to have positively charged regions for favourable

interactions to take place Figure 7 highlights the amino acyl residues that are proposed to interact with 5-methyltetrahydrofolate, according to this model Calculations using interaction potentials produced predicted values for the percentage inhibition of each of the tested compounds These data were then compared with the experimentally determined data (percentage inhibition at 100 lm), and the results are presented in Fig 8 The predicted activities of seven heterocycles (1j, 1k, 1f, 1g, 1a, 2a, 2c) were found to have good cor-relation with the experimentally determined inhibition

Fig 4 Sequence homology of the template sequence of MeTr and the methyltetrahydrofolate-binding domain of rat liver MetS The residues shown in bold indicate conserved homology between the MeTr and MetS proteins Marked with a cross (+) are the residues that have a high degree of similarity so that although the sequence is not identical, the function of the residues is expected to remain the same.

Fig 5 The orientation of hydroxymethylpterin pyrophosphate (HMPP) when bound to dihydropteroate synthase, and the super-imposed structure of methyltetrahydrofolate showing the proposed orientation in the methionine synthase active site HMPP is repre-sented as a stick structure; methyltetrahydrofolate is reprerepre-sented

as a wire structure.

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(percentage inhibition ± 10, Fig 8), including two of

the five most active compounds (i.e the benzimidazoles

1j and 1k) Interestingly, 2b, the most active compound

of this series, was not predicted to have strong inhibitory

activity This finding is consistent with the kinetic

evalu-ation for this inhibitor that suggests a different mode of

action (i.e through noncompetitive inhibition, as shown

in Fig 2) unrelated to direct binding to the methyltetra-hydrofolate site

Overall, the biological evaluation and molecular modelling studies indicate two routes for the develop-ment of the next generation of inhibitors Specifically, the molecules need to be relatively small or not carry bulky substituents in order to enter the active site They require a planar region that can be positioned in the centre of the active site, a nitro functional group and two or three possible hydrogen-bonding groups Further refinement of this model could assist the dis-covery of the next generation of inhibitors for MetS Moreover, the observed noncompetitive inhibition pat-tern, with respect to the methyltetrahydrofolate-bind-ing site of MetS, implies the existence of other bindmethyltetrahydrofolate-bind-ing sites that may also be investigated for the development

of inhibitors, whilst the potential reactivity of the benzothiadiazole ring opens the possibility for design-ing mechanism-based inhibitors Overall, this approach may lead to the identification of compounds with potential therapeutic value, in particular as chemo-therapeutic agents for methionine-dependent cancers in combination with methionine-depleted treatments [33]

As the enzyme and its related metabolites have been involved in many disorders, including cardiovascular disease, neurodegenerative diseases and cancer, potent and specific inhibitors will also be valuable tools for defining the exact role of MetS in the pathophysiology

of these diseases

Experimental procedures

hydroxycobalamin, dimethylsulfoxide, ascorbic acid, phenyl-methanesulfonyl fluoride, Na-p-tosyl-l-lysylchloromethyl ketone, trypsin inhibitor, aprotinin, DEAE-cellulose, and phosphate buffers were purchased from Sigma (Poole, UK) 5-[14C]-methyl]methyltetrahydrofolic acid (barium salt) (56 mCiÆmmol)1) was purchased from Amersham (Little Chalfont, UK) AG1-X8 resin (200–400 mesh chloride form)

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and the Protein Assay kit were from Bio-Rad (Hemel Hemp-stead, UK) Q-Sepharose Fast Flow and Hydroxyapatite were from Pharmacia

HiSafe 3 scintillation cocktail was from Fisher Scientific (Leicester, UK) Amicon ultrafiltration membranes, of

30 kDa, were purchased from

Benzimidazole (1a), 1-methylbenzimidazole (1b), 2,1,3-ben-zothiadiazole (2a), 4-nitro-2,1,3-benzothiadiazole (2b), 4-amino-2,1,3-benzothiadiazole (2c) and 4-methoxyaniline were obtained from Aldrich (Poole, UK); and 1,3-dinitroben-zene was from Avocado (Hewsham, UK) Solvents were

of the highest purity commercially available and were

A

B

C

Fig 6 Snapshot pictures showing the electrostatic surfaces of the

methyltetrahydrofolate-binding domain of rat liver MetS (A) with

5-methyltetrahydrofolate bound showing the open cleft binding side

from a side angle, (B) from a reverse angle, and (C) from above.

Red indicates negatively charged surfaces, and blue indicates

posi-tively charged surfaces.

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purchased from Sigma or BDH (Poole, UK) TLC plates

(sil-ica gel 60 F254) and silica gel (particle size 40–63 lm) for

chromatography were from Merck (Beeston, UK)

Deuterat-ed solvents were from Goss (Glossop, UK) Melting points

were determined using an Electrothermal IA9200 digital melt-ing point apparatus IR spectra were recorded on a Perkin Elmer

Elmer, Seer Green, UK) 1H and 13C NMR spectra were acquired at 270.05 and 67.80 MHz, respectively, on a JEOL GX270 spectrometer (JEOL UK, Welwyn, UK)

assign-ments were made using the DEPT135 experiment Mass spectra were obtained from the EPSRC National Mass Spectrometry Service Centre, University of Wales (Swansea, UK)

Synthesis of the substituted benzimidazoles 1c–k 1,3,5-Trinitrobenzene [34]

1,3-Dinitrobenzene 50 g (0.297 mol) was dissolved in fum-ing nitric acid (130.5 mL) and fumfum-ing sulphuric acid (243.5 mL), then heated under reflux at 150C for 7 days The reaction was cooled slowly to room temperature On addition to ice-cold distilled water, a solid precipitated which was collected by filtration and recrystallized from glacial acetic acid to give 1,3,5-trinitrobenzene (61.01 g, 97%), melting point (m.p.) 118–119C, literature

[34] 1H NMR (CDCl3) d: 9.41 (s

NMR (CDCl3) d: 149.6 (C-NO2), 124.4 (CH-Ar) MS (EI):

213 (M+) IR vmaxÆcm)13104s

13;14 (C-H aromatic), 1624s (C¼C aromatic), 1475m

13;14 (C¼C aromatic), 1544s (N¼O, asym-metric), 1345s (N¼O, symasym-metric), 900s (C-H bend)

Fig 7 Detailed view into the 5-methyltetrahydrofolate-binding pocket of rat liver MetS Atoms within 7 A ˚ of the docked ligand are shown: hydrophobic amino acid residues are coloured bronze, and hydrophilic acid residues are coloured blue Blue lines indicate the putative hydro-gen bonding interactions with ASN100 H-bond lengths are shown on the drawing, deviations from linearity are < 15 for both.

Fig 8 Correlation of experimentally determined inhibition

(percent-age inhibition at 100 l M ) with computer-predicted inhibition, based

on the calculation of interaction potentials.

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3,5-Dinitroanisole [35]

1,3,5-Trinitrobenzene (5 g, 0.023 mol) was dissolved in

methanol (75 mL) with gentle heating To this hot solution,

a hot solution of potassium bicarbonate (0.5 mol, 7.5 g) in

water (30 mL) and methanol (20 mL) was added The

mix-ture was heated at reflux for 2.5 h, cooled to room

tem-perature and the methanol evaporated under reduced

pressure The aqueous residue was extracted with

chloro-form (3· 40 mL), the chloroform extracts combined, dried

over MgSO4and the solvent evaporated The product was

recrystallized from ethanol to give 3,5-dinitroanisole

(3.36 g, 74%), m.p

NMR (CDCl3) d: 8.65 (d

J¼ 2 Hz, 2H, 2-H, 6-H), 4.01 (s, 3H, OCH3) 13C NMR

(dimethylsulfoxide) d: 164.3 (C-1), 152.7 (C-3,5), 118.9

(C-2,6), 114.3 (C-4), 61.07 (CH3) MS (EI): 198 (M+) IR,

vmaxÆcm)1: 3098s (C-H aromatic), 2862w

(C¼C aromatic), 1544s (N¼O, asymmetric), 1345s (N¼O,

symmetric), 1080s (C-O)

2,3,5-Trinitroanisole [36]

3,5-Dinitroanisole (1.5 g, 0.007 mol) was dissolved in

con-centrated sulfuric acid (20 mL) with gentle heating The

solution was placed in an ice bath and fuming nitric acid

(4.2 mL) was added dropwise over a period of 10 min

The mixture was kept on ice for 20 min, with constant

stopped by the addition of distilled water (50–100 mL)

and the product was extracted into ether (3· 40 mL)

The ether extracts were combined, dried over MgSO4and

the solvent evaporated The product was recrystallized

from ethanol to give 2,3,5-trinitroanisole (1.6 g, 96%),

m.p 100–104C, lit 104 C [36] 1

6-H), 4.06 (s, 3H, OCH3) 13C NMR

(dimethylsulfoxide-d6) d: 155.1 (C-1), 152.5 (C-5), 143.9 (C-3), 140.0 (C-2),

116.5 (C-6), 107.4 (C-4), 60.2 (CH3) MS (EI): 243 (M+)

IR vmaxÆcm)1: 3117m (C-H aromatic), 2992w (C-H sp3),

1600m (C¼C aromatic), 1469m (C¼C aromatic), 1046s

(C-O symmetric)

2-Amino-3,5-dinitroanisole [37]

2,3,5-Trinitroanisole (1 g, 0.004 mol) was dissolved in

abso-lute ethanol (54 mL), cooled to 2C, and concentrated

reflux for 3 h, cooled to room temperature and the solvent

evaporated under reduced pressure After isolation by flash

chromatography [chloroform⁄ petroleum ether (70 : 30;

v⁄ v)], the product was recrystallised from ethanol to give

2-amino-3,5-dinitroanisole (0.86 g, 80%), m.p 182–184C,

lit 180C [37] 1H NMR (CDCl3) d: 8.82 (d, J¼ 2 Hz,

1H, 4-H), 9.0–5.0 (br

vmax⁄ cm)1: 3465s (NH asymmetric), 3322s (NH symmetric), 3098w (C-H aromatic), 2992w (C-H sp3), 1600m (C¼C aro-matic), 1456m (C¼C aroaro-matic), 1550s (N¼O asymmetric), 145s (N¼O symmetric), 1059s (C-O)

2-N-Methylamino-3,5-dinitroanisole [37]

2,3,5-Trinitroanisole (1 g, 0.004 mol) was dissolved in tetrahydrofuran (THF) (5 mL) and methylamine in THF

19

(10 mL, 2 m), then the solution was heated in a Young’s tube for 4 h, cooled and the solvent evaporated under reduced pressure The product was isolated by flash chro-matography [diethyl ether⁄ hexane (60 : 40, v ⁄ v)] to give 2-aminomethyl-3,5-dinitroanisole (0.8 g, 89%), 220–222C, lit 230C [37] 1

H NMR (CDCl3) d: 8.76 (d, J¼ 2 Hz, 1H, 4-H), 8.49 (br, 1H, NH), 7.65 (d, J¼ 2 Hz, 1H, 6-H), 3.94 (s, 3H, OCH3), 3.37 (d, J¼ 6 Hz, 3H, NCH3) 13C NMR (CDCl3) d: 149.6 (C-1), 148.4 (C-5), 138.8 (C-3), 137.2 (C-2), 124.3 (C-6), 118.9 (C-4), 57.2 (OCH3), 33.6 (NCH3) MS (EI): 198 (M+)

2,3-Diamino-5-nitroanisole, 3-amino-2-methylamino-5-nitroanisole [37]

The same method was

2-amino-3,5-dinitro-anisole and 2-N-methylamino-3,5-dinitro2-amino-3,5-dinitro-anisole The appro-priate compound (1.0 mmol) was dissolved in methanol (30 mL) and water (2 mL) Ammonium chloride (573 mg,

10 mmol) and ammonium carbonate (350 mg, 3.64 mmol) were added to the solution A hot solution of sodium sulfide (110 mg, 1.41 mmol in 1.5 mL water) was added dropwise over a time-period of 5 min and the solution heated at reflux for 2 h The reaction mixture was allowed

to cool slowly to room temperature, the volatile solvent evaporated under reduced pressure and the product was isolated by flash chromatography, eluted with methanol:

c NH3: chloroform (1 : 1 : 98) 2,3-Diamino-5-nitroanisole: (0.150 mg, 77%), m.p 173–175C, lit.165–167 C [37] 1H NMR (CDCl3) d: 7.41 (d, J¼ 2 Hz, 1H, 4-H), 7.39 (d,

J¼ 2 Hz, 1H, 6-H), 4.05 (s, 2H, NH2), 3.93 (s, 3H,

3-Amino-2-methylamino-5-nitroanisole: (0.181 mg, 80%), m.p 158–160C 1

H NMR (CDCl3) d: 7.32 (d, J¼ 2 Hz, 1H, 4-H), 7.24 (d,

J¼ 2 Hz, 1H, 6-H), 4.1 (br, s, 3H, NH), 3.87 (s, 3H, OCH3), 2.82 (s, 3H, NCH3)

5-Nitro-7-methoxybenzimidazole (1h) and 1-methyl-5-nitro-7-methoxybenzimidazole (1j) [37]

The same method was

2,3-diamino-5-nit-roanisole and 3-amino-2-methylamino-5-nit2,3-diamino-5-nit-roanisole The compound (1.0 mmol) was dissolved in formic acid (5 mL) and heated at reflux for 2 h The reaction was removed

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(s, 3H, OCH3), 3.29 (s, 3H, NCH3) HRMS (ES) (M + H)

208.0717, C9H10N3O3requires 208.0717

5-Amino-7-methoxybenzimidazole (1i) [37] and

5-amino-7-methoxy-N1-methylbenzimidazole (1k) [38]

The same method was

5-nitro-7-methoxy-benzimidazole (1h) and

1-methyl-5-nitro-7-methoxybenzimi-dazole (1j) The compound (0.6 mmol) was dissolved in

ethanol (30 mL) with 2 drops of concentrated HCl, and

10% weight of palladium on a carbon catalyst was added

The system was evacuated and the mixture stirred

vigo-rously under a hydrogen atmosphere until the reaction was

complete (approximately 2–3 h by TLC) The catalyst was

removed by filtration through celite and washed with

copi-ous amounts of ethanol The solvent was evaporated under

reduced pressure and the product was recrystallized from

ethanol and ethyl acetate An alternative method involved

using eight equivalents of ammonium formate as the

hydro-gen source and reacting the mixture, as above, for 2 h in an

evacuated system 1i: (0.08 mg, 82%), m.p 220–223C, lit

216–218C [37].1H NMR (dimethylsulfoxide-d6) d: 8.72 (s,

1H, 2-H), 8.20 (d, J¼ 2 Hz, 1H, 4-H), 7.65 (d, J ¼ 2 Hz,

1H, 6-H), 5.81–6.12 (br, 3H, NH, NH2) 4.07 (s, 3H,

OCH3) 1 k: (0.081 mg, 76%), m.p 180–182, lit 178C

[38].1H NMR (dimethylsulfoxide-d6) d: 8.64 (s, 1H, 2-H),

8.26 (d, J¼ 2 Hz, 1H, 4-H), 7.12 (d, J ¼ 2 Hz, 1H, 6-H),

5.23–5.65 (br, 2H, NH2), 4.01 (s, 3H, OCH3), 3.26 (s, 3H,

NCH3)

5-Nitrobenzimidazole (1c) and 5-aminobenzimidazole

(1d) [39]

These compounds were synthesized, according to the

meth-ods described above, from 2,4 dinitroaniline 1c: (2.5 g,

88%), m.p 203–204C, lit 204–205 C [39] 1

H NMR (dimethylsulfoxide-d6) d: 8.54 (s, 1H, 2-H), 8.51 (d, J¼ 2 Hz,

1H, 4-H), 8.44 (s, 1H, NH), 8.13 (dd

1H, 6-H) 7.09 (d, J¼ 8 Hz, 1H, 7-H) MS (EI): 164(M+

)

1dÆ2HClÆ0.2H2O: (3.1 g, 90%), m.p decomposition

(2· 30 mL) The combined organic extracts were dried over MgSO4 and evaporated under reduced pressure The residue was dissolved in isopropanol, treated with 5 mL of concentrated HCl, evaporated twice from isopropanol then recrystallized from isopropanol-ether to yield a grey solid, 1eÆHCl: (0.50 g, 59%), m.p 202–206C, lit 199–202 C [40] 1H NMR (dimethylsulfoxide-d6) d: 15.01 (br, 2H,

2· NH), 9.43 (d, J ¼ 5 Hz, 1H, 2-H), 7.71 (d, J ¼ 9 Hz, 7-H), 7.23 (d, J¼ 2 Hz, 1H, 4-H), 7.14 (dd, J ¼ 9Hz, J ¼

2 Hz, 1H, 6-H), 3.83 (s, 3H, OCH3) MS (EI) (free base):

148 (M+)

1-Methyl-5-nitro-benzimidazole (1f)

Chlorodinitrobenzene (0.81 g, 4.0 mmol), dissolved in THF

for 12 h at 90C in a Young’s tube The reaction was monitored using TLC with diethyl ether as the eluant The mixture was cooled to room temperature and the solvent evaporated under reduced pressure The resulting diamino compound was then cyclized in formic acid (5 mL) heated

at reflux for 2 h, after which the reaction was cooled to room temperature and toluene (20 mL) and water (1 mL) were added The volatile solvent was evaporated under reduced pressure and the residue was poured into water (30 mL) and extracted with ethyl acetate (3· 30 mL) The

(20 mL), dried over MgSO4and evaporated under reduced pressure The product was recrystallized from ethyl acetate (0.23 g, 32%), m.p 213–215C, lit 209–211 C [41] 1

H NMR (dimethylsulfoxide-d6) d: 8.73 (d, J¼ 2 Hz, 1H, 4-H), 8.25 (d, J¼ 9 Hz, 2H, 6-H), 8.04 (s, 1H, 2-H), 7.44 (d, J¼ 9 Hz, 1H, 7-H), 3.92 (s, 3H, NCH3) MS (EI):

178 (M+)

1-Methyl-5-amino-benzimidazoleÆ2HClÆ0.2H2O (1g)

1-Methyl-5-nitro-benzimidazole (1f) (0.097 g, 0.548 mmol) was dissolved in ethanol (20 mL), 10% palladium on carbon

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